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Planet X-Philes

Hypergeophilic & Hypergeotolerant Organisms


Introduction

“Hypergeophilic prokaryotes” and “hypergeotolerant prokaryotes” are defined here as prokaryotes with one or more specific “hypergeotolerances”, or adaptations/exaptations to enduring or even thriving optimally in planetary conditions, such as extreme heat, extreme pressure, high acidity, high alkalinity/basicity, and high salinity. Therefore, included in this category are thermophiles, barophiles, acidophiles, alkaliphiles, and halophiles. These are in contrast to astrotolerant organisms, or creatures capable of surviving interplanetary extremes.

In addition to other sources cited, much of this information was compiled with the help of resources such as the The List of Prokaryotic names with Standing in Nomenclature (LPSN), and NCBI’s Taxonomy Browser.




Taxonomic/Phylogenetic Overview of Hypergeophilic & Hypergeotolerant Genera

“Hypergeophilic genera” and “hypergeotolerant genera” refer here to genera known to include hypergeophilic species and/or species with hypergeotolerances.

Genera containing thermophilic and hyperthermotolerant organisms are marked with an asterisk (*)

Genera containing barophilic and hyperbarotolerant organisms are marked with two asterisks (**)

Genera containing acidophilic and hyperacidotolerant organisms are marked with a dagger (†)

Genera containing alkaliphilic and hyperalkalitolerant organisms are marked with two daggers (††)

Genera containing thermoacidophilic and hyperthermoacidotolerant organisms are marked with an asterisk and a dagger (*†)

Genera containing halophilic and hyperhalotolerant organisms are marked with a double dagger (‡)

Bacteria marked with an asterisk and a degree sign (*°) are endospore-forming polyextremotolerant organisms. Bacterial endospores tend to give the bacteria capable of producing them polyextremotolerance. Endospores are known for being able to survive extremely high doses of radiation, vacuum dessication, extreme heat and cold, lack of nutrients, and in some cases even the passage of hundreds of millions of years (for example, Virgibacillus marismortui (formerly Bacillus marismortui and Bacillus strain 2-9-3) was revived by scientists from 250-million-year-old spores (Vreeland, 2000); for comparison, the first dinosaurs began appearing about 220 MYA — 30 million years later). Although these endospore-forming bacteria can survive a variety of harsh conditions, endospore-formation is not done for reproduction, as with fungal spore formation, but is instead a last-ditch effort to survive conditions the bacterium would not otherwise be able to, which means these critters aren’t true polyextremophiles, because they don’t (necessarily) actually grow better in extreme conditions, and in their endospore form don’t even grow at all.

In addition to other sources cited, much of this information was compiled with the help of NCBI’s Taxonomy Browser.


Domain: Archaea
Phylum: Euryarchaeota
Class: Archaeoglobi
Order: Archaeoglobales
Family: Archaeoglobaceae
Genus: Archaeoglobus *
Class: Stenosarchaea
Order: Candidatus Nanohaloarchaeota
Family: Nanohaloarchaea
Genus: Haloredivivus †‡
Genus: Nanosalina *†‡
Genus: Nanosalinarum †‡
Class: Halobacteria
Order: Halobacteriales
Family: Haloarculaceae
Genus: Haloarcula
Genus: Halorhabdus
Genus: Halorientalis
Genus: Halosimplex
Genus: Natronomonas ††‡
Family: Halobacteriaceae
Genus: Haladaptatus
Genus: Halalkalicoccus ††‡
Genus: Halarchaeum †‡
Genus: Haloalcalophilum ††‡
Genus: Halobacterium
Genus: Halomarina
Genus: Halorussus
Genus: Natronoarchaeum
Genus: Salarchaeum
Family: Halococceae
Genus: Halococcus
Order: Haloferacales
Family: Haloferacaceae
Genus: Halobellus
Genus: Haloferax
Genus: Halopelagius
Genus: Haloplanus
Genus: Haloquadratum
Family: Halorubraceae
Genus: Halobaculum
Genus: Halogeometricum
Genus: Halogranum
Genus: Halolamina
Genus: Halonotius
Genus: Halorubrum ††‡
Order: Natrialbales
Family: Natrialbaceae
Genus: Halobiforma
Genus: Halopiger
Genus: Halostagnicola
Genus: Haloterrigena
Genus: Halovivax
Genus: Natrarchaeobaculum
Genus: Natrialba ††‡+
Genus: Natrinema
Genus: Natronobacterium ††‡
Genus: Natronococcus ††‡
Genus: Natronolimnohabitans
Genus: Natronolimnobius
Genus: Natronorubrum
Class: Methanococci
Order: Methacoccales
Family: Methanococcaceae
Genus: Methanocaldococcus ***
Class: Methanopyri
Order: Methanopyrales
Family: Methanopyraceae
Genus: Methanopyrus *
Class: Thermococci
Order: Thermococcales
Family: Thermococcaceae
Genus: Pyrococcus ***
Genus: Thermococcus ***†††
Class: Thermoplasmata
Order: Methanomassiliicoccales
Family: Methanomassiliicoccaceae
Genus: Methanomassiliicoccus
Order: Thermoplasmatales
Family: Cuniculiplasmaceae
Genus: Cuniculiplasma
Family: Ferroplasmaceae
Genus: Acidiplasma
Genus: Ferroplasma
Family: Picrophilaceae
Genus: Picrophilus
Family: Incertae sedis
Genus: Thermogymnomonas
Superphylum: DPANN group
Phylum: Candidatus Diapherotrites
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Iainarchaeum
Phylum: Aenigarchaeota
Class: (unnamed?)
Order: Aenigmarchaeales
Family: (unnamed?)
Genus: Aenigmarchaeum
Phylum: Nanoarchaeota
Class: (unnamed?)
Order: Nanoarchaeales
Family: Nanoarchaeaceae
Genus: Nanoarchaeum *†
Family: Nanopusillaceae
Genus: Nanobsidianus *†
Genus: Nanopusillus *†
Phylum: Incertae sedis
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Mancarchaeum †
Phylum: Candidatus Micrarchaeota
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Micrarchaeum †
Phylum: Candidatus Parvarchaeota
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Parvarchaeum †
Kingdom: Proteoarchaeota
Superphylum: TACK group
Phylum: Thaumarchaeota
Class: Nitrososphaeria
Order: Nitrosopumilales
Family: Nitrosopumilaceae
Genus: Candidatus Nitrosoarchaeum †
Genus: Nitrosopumilus †
Order: Nitrososphaerales
Family: Nitrososphaeraceae
Genus: Candidatus Nitrocosmicus †
Genus: Candidatus Nitrososphaera †
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Nitrosopelagicus †
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Nitrosotalea †
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Nitrosotenius †
Phylum: Crenarchaeota
Class: Thermoprotei
Order: Desulfurococcales
Family: Desulfurococcaceae
Genus: Aeropyrum *
Genus: Candidatus Nanopusillus †
Family: Pyrodictiaceae
Genus: Geogemma *
Genus: Pyrolobus *
Order: Sulfolobales
Family: Sulfobaceae
Genus: Acidianus *†
Genus: Metallosphaera *†
Genus: Stygiolobus *†
Genus: Sulfolobus *†
Genus: Sulfurisphaera
Genus: Sulfurococcus
Class: Incertae sedis
Order: Cenarchaeales
Family: Cenarchaeaceae
Genus: Cenarchaeum

Domain: Bacteria
Phylum: Acidobacteria
Class: Acidobacteriia
Order: Acidobacteriales
Family: Acidobacteriaceae
Genus: Acidobacterium
Genus: Bryocella
Genus: Telmatobacter
Phylum: Aquificae
Class: Aquificidae
Order: Aquificales
Family: Aquifaceae
Genus: Aquifex *
Genus: Hydrogenobacter *†
Genus: Hydrogenobaculum *†
Phylum: Desulfobacteriota
Class: Thermodesulfobacteria
Order: Thermodesulfobacteriales
Family: Thermodesulfobacteriaceae
Genus: Geothermobacterium *
Phylum: Nitrospirae
Class: Nitrospira
Order: Nitrospirales
Family: Nitrospiraceae
Genus: Leptospirillum
Phylum: Proteobacteria
Class: Acidithiobacilla
Order: Acidithiobacillales
Family: Acidithiobacillaceae
Genus: Acidithiobacillus
Class: Alphaproteobacteria
Order: Rhodospirallales
Family: Acetobacteraceae
Genus: Acetobacter
Genus: Acidiphilium
Genus: Acidocella
Genus: Acidomonas
Class: Betaproteobacteria
Order: Burkholderiales
Family: Comamonadaceae
Genus: Thiomonas
Order: Nitrosomonadales
Family: Gallionellaceae
Genus: Gallionella
Family: Thiobacillaceae
Genus: Thiobacillus
Class: Gammaproteobacteria
Order: Chromatiales
Family: Ectothiorhodospiraceae
Genus: Halorhodospira ††‡
Genus: Thiohalospira ††‡
Clade: Delta/Epsilon subdivisions
Class: Deltaproteobacteria
Order: Desulfobacterales
Family: Desulfobacteraceae
Genus: Desulfobacter
Genus: Desulfosarcina
Family: Desulfobulbaceae
Genus: Desulfobulbus
Order: Desulfovibrionales
Family: Desulfomicrobiaceae
Genus: Desulfomicrobium
Family: Desulfovibrionaceae
Genus: Desulfovibrio
Class: Epsilonproteobacteria
Order: Campylobacterales
Family: Helicobacteraceae
Genus: Helicobacter
Phylum: Thermotogae
Class: ???
Order: Thermotogales
Family: Thermotogaceae
Genus: Thermotoga *
Kingdom: Terrabacteria
Phylum: Actinobacteria
Class: Acidimicrobiia
Order: Acidimicrobiales
Family: Acidimicrobiaceae
Genus: Acidiferrimicrobium
Genus: Acidimicrobium
Genus: Ferrimicrobium
Phylum: Firmicutes
Class: Bacilli
Order: Bacillales
Family: Alicyclobacillaceae
Genus: Alicyclobacillus *†°
Family: Bacillaceae
Genus: Bacillus
Genus: Virgibacillus *°‡
Class: Clostridia
Order: Eubacteriales
Family: Clostridiaceae
Genus: Anoxynatronum ††
Genus: Clostridium
Family: Peptococcaceae
Genus: Desulfotomaculum
Family: Incertae sedis
Genus: Sulfobacillus *°†



Alphabetical List of Hypergeophilic & Hypergeotolerant Genera

Genera containing thermophilic and hyperthermotolerant organisms are marked with an asterisk (*)

Genera containing barophilic and hyperbarotolerant organisms are marked with two asterisks (**)

Genera containing acidophilic and hyperacidotolerant organisms are marked with a dagger (†)

Genera containing alkaliphilic and hyperalkalitolerant organisms are marked with two daggers (††)

Genera containing halophilic and hyperhalotolerant organisms are marked with a double dagger (‡)

In addition to other sources cited, much of this information was compiled with the help of NCBI’s Taxonomy Browser.

Described genera
Genus Source
Acetobacter ()
Acidiferrimicrobium Gonzalez &al., 2020
Acidimicrobium
Acidianus ()
Acidiphilium ()
Acidiplasma Zhang &al., 2016
Acidithiobacillus (Fashola &al., 2015)
Acidobacterium
Acidocella Johnson, 1998
Acidomonas Johnson, 1998
Aenigmarchaeum Golyshina &al., 2017a
* Aeropyrum Sako &al., 1996; Baker &al., 2006
*†° Alicyclobacillus Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
*†*† Anoxynatronum ?
* Aquifex ?
* Archaeoglobus Klenk &al., 1997
Bacillus ?
Bryocella elongata ()
Cenarchaeum Herbold &al., 2017
Clostridium ?
Cuniculiplasma Golyshina &al. 2016
Desulfobacter Fashola &al., 2015
Desulfobulbus Fashola &al., 2015
Desulfomicrobium sp. ?
Desulfosarcina Fashola &al., 2015
Desulfotomaculum Fashola &al., 2015
Desulfovibrio Fashola &al., 2015
** Exiguobacterium ?
Ferroplasma Edwards &al., 2000, Dopson &al. 2003; Zhang &al., 2016
Gallionella Ehrenberg, 1838
* Geogemma Kashefi & Lovley, 2003
* Geothermobacterium ?
Haladaptatus Savage &al., 2007
††‡ Halalkalicoccus Xue &al., 2005
†‡ Halarchaeum ()
†‡ Haloalcalophilum ()
Haloarcula (Torreblanca &al.., 1986) Oren &al., 2009
Halobacterium Elazari-Volcani, 1957
Halobaculum ?
Halobellus ?
Halobiforma Hezayen &al., 2002
Halococcus Schoop, 1935
Haloferax ?
Halogeometricum ?
Halogranum ?
Halolamina ?
Halomarina ?
Halomicrobium ?
Halonotius ?
Halopelagius ?
Halopiger ?
Haloplanus ?
Haloquadratum ?
†‡ Candidatus Haloredivivus Baker &al., 2006; Golyshina &al., 2017a
Halorhabdus ?
††‡ Halorhodospira ?
Halorientalis ?
††‡ Halorubrum Oren, 2002
Halorussus ?
Halosimplex ?
Halostagnicola ?
Haloterrigena ?
Halovivax ?
Helicobacter
*† Hydrogenobacter
*† Hydrogenobaculum Fashola &al., 2015
Iainarchaeum Rinke, &al., 2003
Leptospirillum Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
Candidatus Mancarchaeum Baker &al., 2006; Golyshina &al., 2017a
*† Metallosphaera Zhang &al., 2016
*** Methanocaldocococcus Brock &al., 1972
* Methanopyrus Kurr &al., 1992
Candidatus Micrarchaeum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017a
*† Candidatus Nanoarchaeum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017a
Nanobsidianus
Nanopusillus
*†‡ Nanosalina Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017???
†‡ Nanosalinarum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017???
††‡+ Natrialba Oren, 2002
Natrinema
Natrarchaeobaculum
Natronoarchaeum
††‡ Natronobacterium Oren, 2002
††‡ Natronococcus
Natronolimnobius
††‡ Natronomonas
Natronorubrum
Nitrosoarchaeum Herbold &al., 2017
Nitrosocosmicus Herbold &al., 2017
Nitrosopelagicus Herbold &al., 2017
Nitrosopumilus Lehtovirta-Morley &al., 2011 & 2014; Herbold &al., 2017
Nitrososphaera Herbold &al., 2017
Nitrosotalea Herbold &al., 2017
Nitrosotenuis Herbold &al., 2017
Parvarchaeum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017???
Picrophilus (Baker-Austin & Dopson, 2007)
*** Pyrococcus Erauso &al., 1993
* Pyrolobus Bloch &al., 1997
Salarchaeum ?
* Stygiolobus Segerer &al., 1991
*† Sulfolobus Brock &al., 1972
*°† Sulfobacillus
Sulfurisphaera Zhang &al., 2016
Sulfurococcus Johnson, 1998
***††† Thermococcus ?
Telmatobacter ()
Thermogymnomonas ?
Thermoplasma Johnson, 1998; Zhang &al., 2016
Thiobacillus Johnson, 1998
††‡ Thiohalospira ?
Thiomonas Johnson, 1998
*°‡ Virgibacillus



List of Hypergeophilic & Hypergeotolerant Species

In addition to other sources cited, much of this information was compiled with the help of resources such as the The List of Prokaryotic names with Standing in Nomenclature (LPSN), and NCBI’s Taxonomy Browser.

Note: This list is still under construction and has definitely not yet been double-checked for accuracy! This situation will be remedied as quickly as possible.

Described Species
Genus & Species Source
Acetobacter aceti Johnson, 1998
Acidianus ambivalens Johnson, 1998; Zhang &al., 2016
Acidianus brierleyi Johnson, 1998; Edwards &al., 2000; Fashola &al., 2015; Zhang &al., 2016
Acidianus copahuensis Zhang &al., 2016
Acidianus infernus Johnson, 1998
Acidianus manzaensis Ding &al. 2011; Zhang &al., 2016
Acidianus pozzuoliensis
*† Acidianus sulfidivorans Plumb &al. 2007; Zhang &al., 2016
Acidianus tengchongensis He &al. 2004; Zhang &al., 2016
Acidianus sp. DSM 29099 (formerly sp. RZ1) Zhang &al., 2016
Acidianus convivator ()
Acidiferrimicrobium australe Gonzalez &al., 2020
Acidimicrobium ferroxidans Johnson, 1998; Fashola &al., 2015
Acidiphilium acidophilum (more species?)
(Thiobacillus acidophilus)
Johnson, 1998
Acidiphilium multivorum (Dopson 2003)
Acidiplasma aeolicum Zhang &al., 2016
Acidiplasma cupricumulans Zhang &al., 2016
Acidithiobacillus albertensis
(Thiobacillus albertis)
Johnson, 1998; Fashola &al., 2015
Acidithiobacillus caldus
(Thiobacillus caldus)
Johnson, 1998;
Fashola &al., 2015; Zhang &al., 2016
Acidithiobacillus ferridurans ()
Acidithiobacillus ferriphilus ()
Acidithiobacillus ferrivorans (Fashola &al., 2015)
Acidithiobacillus ferroxidans (formerly Thiobacillus ferrooxidans) Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
Acidithiobacillus thioxidans (formerly Thiobacillus thioxidans) Johnson, 1998; Fashola &al., 2015
Acidobacterium capsulatum Johnson, 1998; Fashola &al., 2015
Acidocella aluminiidurans Kimoto &al., 2010; Johnson, 1998
Acidocella aminolytica (Kishimoto &al., 1994) Kishimoto &al., 1996; Johnson, 1998
Acidocella aquatica Okamoto &al., 2017; Johnson, 1998
Acidocella aminolytica (Wichlacz &al., 1986) Kishimoto &al., 1996; Johnson, 1998
Acidomonas methanolica Johnson, 1998
Candidatus Aenigmarchaeum subterraneum Baker &al., 2006; Golyshina &al., 2017a
* Aeropyrum pernix Sako &al., 1996; Baker &al., 2006
*†° Alicyclobacillus acidiphilus Matsubara &al., 2002
*†° Alicyclobacillus acidocaldarius (Darland & Brock, 1971) Wisotskey &al., 1992
*†° Alicyclobacillus acidoterrestris (Deinhard &al., 1988) Wisotskey &al., 1992
*†° Alicyclobacillus aeris Guo &al., 2009
*†° Alicyclobacillus cellulosilyticus Kusube &al., 2014
*†° Alicyclobacillus contaminans Goto &al., 2007
*†° Alicyclobacillus cycloheptanicus Matsubara &al., 2002
*†° Alicyclobacillus dauci Nakano &al., 2015
*†° Alicyclobacillus disulfidooxidans (Dufresne &al., 1996) Karavaiko &al., 2005
*†° Alicyclobacillus fastidiosus Goto &al., 2007
*†° Alicyclobacillus ferrooxydans Jiang &al., 2008
*†° Alicyclobacillus fodiniaquatilis Zhang &al., 2015a
*†° Alicyclobacillus herbarius Goto &al., 2002
*†° Alicyclobacillus hesperidum Albuquerque &al., 2000
*†° Alicyclobacillus kakegawensis Goto &al., 2002
*†° Alicyclobacillus macrosporangiidus Goto &al., 2007
*†° Alicyclobacillus montanus Lopez &al., 2018
*†° Alicyclobacillus pomorum Goto &al., 2003
*†° Alicyclobacillus sacchari Goto &al., 2007
*†° Alicyclobacillus sendaiensis Tsuruoka &al., 2003
*†° Alicyclobacillus shizuokensis Goto &al., 2007
*†° Alicyclobacillus tengchongensis Kim &al., 2016
*†° Alicyclobacillus thermosulfidooxidans
(Sulfobacillus thermosulfidooxidans)
Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
*†° Alicyclobacillus tolerans Karavaiko &al., 2005
*†° Alicyclobacillus vulcanalis Simbahan &al., 2004
* Archaeoglobus fulgidus Klenk &al., 1997
* Archaeoglobus infectus Mori &al., 2008
* Archaeoglobus lithotrophicus ?
* Archaeoglobus profundis Burggraf &al. 1990
* Archaeoglobus veneficus Huber &al. 1998
Bacillus acidicola Albert &al. 2005
Bacillus aerius Shivaji &al., 2014
Bacillus aerophilus Shivaji &al., 2006
Bacillus albus Liu &al., 2017
Bacillus altitudinis Shivaji &al., 2006
Bacillus alveayuensis Bae &al., 2005
Bacillus amyloliquefaciens (Fukumoto, 1943) Priest &al., 1987
Bacillus anthracis (Anthrax) Cohn, 1872
Bacillus aquiflavi Xie &al., 2020
Bacillus atrophaeus Nakamura, 1989
Bacillus australimaris Liu &al., 2016
Bacillus badius Batchelor, 1919
Bacillus benzoevorans Pichinoty &al., 1987
Bacillus cabrialesii de Los Santos Villalobos &al., 2019
Bacillus canaveralius Newcombe &al., 2009
Bacillus capparidis Wang &al., 2017
Bacillus cereus Frankland & Frankland, 1887
Bacillus chungangensis Cho &al., 2010
Bacillus cahuilensisereus Cerritos &al., 2008
Bacillus cytotoxicus Guinebretiere &al., 2013
Bacillus decisifrondis Zhang &al., 2007
Bacillus ectoiniformans Zhu &al., 2016
Bacillus enclensis Dastager &al., 2014
Bacillus fengquiuensis Zhao &al., 2014
Bacillus fungorum Liu &al., 2020
Bacillus glycinifermentans Kim &al., 2015
Bacillus gobiensisectoiniformans Liu &al., 2016
Bacillus halotolerans (Delaporte & Sasson, 1967) Tindall &al., 2017
Bacillus heynesii Dunlap &al., 2017
Bacillus horti Yumoto &al., 1998
Bacillus inaquosorum (Rooney &al., 2009) Dunlap &al., 2020
Bacillus infantis Ko &al., 2006
Bacillus infernus Boone &al., 1995
Bacillus isabeliae Albuquerque &al., 2008
Bacillus kexueae Sun &al., 2018
Bacillus licheniformis (Weigmann, 1898) Chester, 1901
Bacillus luti Liu &al., 2017
Bacillus manusensis Sun &al., 2018
Bacillus marinisedimentorum Guo &al., 2018
Bacillus mesophilus Manickam &al., 2014
Bacillus methanolicus Arfman &al., 1992
Bacillus mobilis Liu &al., 2017
Bacillus mojavensis Roberts &al., 1994
Bacillus mycoides Flugge &al., 1886
Bacillus nakamurai Dunlap &al., 2016
Bacillus ndiopicus Lo &al., 2017
Bacillus nitratireducens Guo &al., 2016
Bacillus oleivorans Azmatunnisa &al., 2015
Bacillus pacificus Liu &al., 2017
Bacillus pakistanensis Roohi &al., 2014
Bacillus paralicheniformis Dunlap &al., 2015
Bacillus paramycoides Liu &al., 2017
Bacillus paranthracis Liu &al., 2017
Bacillus pervagus Kosowski &al., 2014
Bacillus priscicola Lee &al., 2016
Bacillus proteolyticus Liu &al., 2017
Bacillus pseudomycoides Nakamura, 2018
Bacillus pumilus Meyer & Gottheil, 1901
Bacillus safensis Satomi &al., 2006
Bacillus salacetis Daroonpunt &al., 2019
Bacillus salinus Kang &al., 2020
Bacillus salitolerans Zhang &al., 2015b
Bacillus seohaeanensis Lee &al., 2006
Bacillus siamensis Sumpavapol &al., 2010
Bacillus shivajii Kumar &al., 2018
Bacillus smithii Nakamura &al., 1988
Bacillus solimangrovi Lee &al., 2014
Bacillus songklensis Kang &al., 2013
Bacillus sonoresnis Palmisano &al., 2001
Bacillus spizizenii (Nakamura &al., 1999) Dunlap &al., 2020
Bacillus spongiae Lee &al., 2019
Bacillus stercoris (Adelskov & Patel, 2017) Dunlap &al., 2020
Bacillus stratosphericus Shivaji &al., 2006
Bacillus subtilis (Ehrenberg 1835) Cohn 1872
Bacillus swezeyi Dunlap &al., 2017
Bacillus taeanensis Lim &al., 2006
Bacillus tamaricis Zhang &al., 2018
Bacillus tequilensis Gatson &al., 2006
Bacillus thermocloacae Demharter & Hensel, 1989
Bacillus thermotolerans Yang &al., 2013
Bacillus thuringiensis Burlinger, 1915
Bacillus tianshenii Jiang &al., 2014
Bacillus toyonensis Jimenez &al., 2014
Bacillus tropicus Liu &al., 2017
Bacillus vallismortis Roberts &al., 1996
Bacillus velezensis Ruis-Garcia &al., 2005
Bacillus wiedmannii Miller &al., 2016
Bacillus wudalienchiensis Liu &al., 2017
Bacillus xiamenensis Lai &al., 2014
Bacillus xiapuensis Liu &al., 2019
Bacillus zhangzhouensis Liu &al., 2016
Bacillus sp. Schuerger & Nicholson, 2016
Bryocella elongata ()
Candidatus Cenarchaeum symbiosum A Herbold &al., 2017
Clostridium aceticum Wieringa, 1940
Clostridium acetireducens Orlygsson &al., 1996
Clostridium acetobutylicum McCoy &al., 1926
Clostridium acidisoli Kuhner &al., 2000
Clostridium aestuarii Kim &al., 2007
Clostridium akagii Kuhner &al., 2000
Clostridium algidicarnis Lawson &al., 1995
Clostridium algifaecis Wu &al., 2014
Clostridium algoriphilum Shcherbakova &al., 2010
Clostridium amazonense O’Neal &al., 2015
Clostridium aminophilum Paster &al., 1993
Clostridium amylolyticum Song & Dong, 2008
Clostridium arbusti Jung &al., 2010
Clostridium arcticum Jordan & McNicol, 1979
Clostridium argentinense Suen &al., 1988
Clostridium atrophaeus Nakamura, 1989
Clostridium aurantibutyricum Hellinger, 1944
Clostridium baratii Prevot, 1938
Clostridium beihaiense Dong &al., 2018
Clostridium beijerinckii Donker, 1926
Clostridium bornimense Hahnke &al., 2014
Clostridium botulinum van Ermengem, 1896
Clostridium bovifaecis Zhu &al., 2018
Clostridium bowmanii Spring &al., 2003
Clostridium budayi Le Blaye, 1914
Clostridium butyricum Prazmowski, 1880
Clostridium cadaveris Klein, 1899
Clostridium carboxidivorans Liou &al., 2005
Clostridium carnis Klein, 1904
Clostridium cavendishii Bowman &al., 2010
Clostridium celatum Hauschild & Holdeman, 1974
Clostridium cellulosi He &al., 1991
Clostridium cellulovorans Sleat &al., 1985
Clostridium chartatabidum Kelly &al., 1996
Clostridium chauvoei Arloing &al., 1887
Clostridium chromiireducens Inglett, 2011
Clostridium clostridioforme Burri & Ankersmit, 1906
Clostridium cochlearium Douglas, 1919
Clostridium cocleatum Kaneuchi &al., 1979
Clostridium colicanis Greetham &al., 2003
Clostridium colinum Berkhoff &al., 1974
Clostridium collagenovorans Jain & Zeikus, 1988
Clostridium combesii Prevot & Laplanche, 1947
Clostridium composti Shin &al., 2018
Clostridium cylindrosporum Barker & Beck, 1942
Clostridium disporicum Horn, 1987
Clostridium drakei Liou &al., 2005
Clostridium estertheticum Collins &al., 1993
Clostridium fallax Weinberg & Seguin, 1915
Clostridium felsineum Carbone & Tombolato, 1917
Clostridium fermenticellae Xu &al., 2019
Clostridium fessum Seo &al., 2020
Clostridium fimetarium Kotsyurbenko &al., 1997
Clostridium formicaceticum Andreesen &al., 1970
Clostridium frigidicarnus Broda &al., 1999
Clostridium frigoris Spring &al., 2003
Clostridium ganghwense Kim &al., 2006
Clostridium gasigenes Broda &al., 2000
Clostridium granti Mountfort &al., 1996
Clostridium guangxiense Zhao &al., 2017
Clostridium haemolyticum Hall, 1929
Clostridium halophilum Fendrich &al., 1991
Clostridium herbivorans Varel &al., 1995
Clostridium hiranonis Kitahara &al., 2001
Clostridium histolyticum
(not very thermotolerant; OGT=60°C)
Weinberg & Seguin, 1916
Clostridium homopropionicum Dorner & Schink, 1991
Clostridium huakuii Ruan &al., 2014
Clostridium hydrogeniformans Bowman &al., 2010
Clostridium hylemonae Kitahara &al., 2000
Clostridium indicum
(extremotolerant mesophile, OGT=37°C)
Gundawar &al., 2019
Clostridium innocuum Smith & King, 1962
Clostridium intestinale Lee &al., 1989
Clostridium irregulare Choukevitch, 1911
Clostridium isatidis Padden &al., 1999
Clostridium jeddahense Lagier &al., 2016
Clostridium kluyveri Barker & Taha, 1942
Clostridium lacusfryxellense Spring &al., 2003
Clostridium leptum Moore &al., 1976
Clostridium liquoris Yin &al., 2016
Clostridium ljungdahlii Tanner &al., 1993
Clostridium lundense Cirne &al., 2006
Clostridium luticellarii Wang &al., 2015
Clostridium magnum Schink, 1984
Clostridium malenominatum Weinberg &al., 1937
Clostridium methoxybenzovorans Mechichi &al., 1999
Clostridium methylpentosum Himelbloom & Canale-Parola, 1989
Clostridium moniliforme Repaci, 1910
Clostridium neonatale Bernard &al., 2018
Clostridium neuense Zhao &al., 2017
Clostridium nexile Holdeman & Moore, 1974
Clostridium nitritogenes Prevot, 1940
Clostridium nitrophenolicum Suresh &al., 2007
Clostridium novyi Migula, 1900
Clostridium oceanicum Smith, 1970
Clostridium oryzae Horino &al., 2015
Clostridium pabulibutyricum Kobayashi &al., 2017
Clostridium paradoxum Li &al., 1993
Clostridium paraputrificum Bienstock, 1906
Clostridium pascui Wilde &al., 1997
Clostridium pasteurianum Winogradsky, 1985
Clostridium peptidovorans Mechichi &al., 2000
Clostridium perfringens
(formerly C. welchii)
Veillon & Zuber, 1898
Clostridium phytofermentans Warnick &al., 2002
Clostridium piliforme Tyzzer, 1917
Clostridium polyendosporum Duda &al., 1996
Clostridium polysaccharolyticum van Glyswyk, 1981
Clostridium populeti Sleat & Mah, 1985
Clostridium prolinivorans Huang &al., 2020
Clostridium psychrophilum Spring &al., 2003
Clostridium punense Lanjekar &al., 2015
Clostridium puniceam Lund &al., 1981
Clostridium putrefaciens McBryde, 1911
Clostridium quinii Svensson &al., 1995
Clostridium ramosum Veillon & Zuber, 1898
Clostridium roseum McCoy & McClung, 1935
Clostridium saccharobutylicum Kleis &al., 2001
Clostridium saccharogumia Clavel &al., 2007
Clostridium saccharoperbutylacetonicum Kleis &al., 2001
Clostridium sardiniense
(aka Clostridium absonum
Prevot, 1938
Clostridium sartagoforme Partansky & Henry, 1935
Clostridium saudiense Angelakis &al., 2016
Clostridium scidens Morris &al., 1985
Clostridium senagelense Mishra &al., 2015
Clostridium septicum Macé, 1889
Clostridium spiroforme Kaneuchi &al., 1979
Clostridium sporogenes Metchnikoff, 1908
Clostridium sporosphaeroides Soriano & Soriano, 1948
Clostridium subterminale Hall & Whitehead, 1927
Clostridium sulfidigenes Sallam & Steinbuchel, 2009
Clostridium swellfunianum Liu &al., 2015
Clostridium symbiosum Stevens, 1956
Clostridium tagluense Suetin &al., 2009
Clostridium tarantellae (Udey &al., 1977) Lawson & Rainey, 2016
Clostridium tepidiprofundi Slobodkina &al., 2008
Clostridium tepidum Dobritsa &al., 2017
Clostridium tertium (Henry, 1917) Bergey &al., 1923
Clostridium tetani (Flugge, 1886) Bergey &al., 1923
Clostridium tetanomorphum (Bulloch &al., 1919) Wilde &al., 1989
Clostridium thermarum Liu &al., 2021
Clostridium thermoalcaliphilum Li &al., 1994
Clostridium thermobutyricum Weigel &al., 1989
Clostridium thermopalmarium Soh &al., 1991
Clostridium thermopapyrolyticum Mendez &al., 1991
Clostridium thermosaccharolyticum McClung, 1935
Clostridium thermosuccinogenes Drent &al., 1995
Clostridium thermosulfurigenes Schink & Zeikus, 1983
Clostridium thiosulfatireducens Hernandez-Euginio &al., 2002
Clostridium tyrobutyricum van Beynum & Pette, 1935
Clostridium uliginosum Matthies &al., 2001
Clostridium villosum Love &al., 1979
Clostridium vincentii Mountfort &al., 1997
Clostridium viride Buckel &al., 1995
Clostridium vulturis Paek &al., 2015
Cuniculiplasma divulgatum Golyshina &al. 2016
Desulfobacter sp. Fashola &al., 2015
Desulfobulbus sp. Fashola &al., 2015
Desulfomicrobium sp. Fashola &al., 2015
Desulfosarcina sp. Fashola &al., 2015
Desulfotomaculum sp. Fashola &al., 2015
** Exiguobacterium sp. Schuerger & Nicholson, 2016
Desulfovibrio sp. Fashola &al., 2015
Ferrimicrobium acidiphilum Johnson, 1998
Ferrimicrobium acidiphilus Fashola &al., 2015
Ferroplasma acidarmanus Edwards &al., 2000, Dopson &al.2003; Zhang &al., 2016
Ferroplasma acidiphilum Fashola &al., 2015; Zhang &al., 2016; Golyshina &al., 2017b
Ferroplasma cyprexacervatum ()
Ferroplasma thermophilum Zhang &al., 2016
Gallionella ferruginea Ehrenberg, 1838; Fashola &al., 2015
* Geogemma barossi
(Pyrolobus fumarii, strain 121)
Kashefi & Lovley, 2003
* Geothermobacterium ferrireducens ?
Haladaptatus cibarius ()
Haladaptatus litoreus ()
Haladaptatus pallidirubidus ()
Haladaptatus paucihalophilus ()
††‡ Halalkalicoccus jeotgali ()
††‡ Halalkalicoccus tibetensis ()
†‡ Halarchaeum acidiphilum ()
Haloalcalophilum atacamensis Oren & Ventosa, 2000
Haloarcula amylolytica ?
Haloarcula argentinensis ?
Haloarcula hispanica ?
Haloarcula japonica ?
Haloarcula marismortui ?
Haloarcula quadrata ?
Haloarcula vallismortis ?
Halobacterium jilantaiense ?
Halobacterium noricense ?
Halobacterium salinarum ?
Halobacterium piscisalsi ?
Halobaculum gomorrense Oren &al., 1995
Halobaculum magnesiipjilum Shimoshige &al., 2013
Halobaculum roseum Chen &al., 2017
Halobaculum saliterrae Myers & King, 2020
Halobellus captivus Chen &al., 2020
Halobellus clavatus Cui &al., 2011
Halobellus inordinatus Qiu &al., 2013
Halobellus Iimi Cui &al., 2012
Halobellus litoreus Zhao &al., 2014
Halobellus ramosii Perez-Davo &al., 2015
Halobellus rarus Zhang &al., 2014
Halobellus refus Cha &al., 2014
Halobellus salinus Cui &al., 2012
Halobiforma haloterrestris ?
Halobiforma lacisalsi ?
Halobiforma nitratireducens ?
Halococcus agarilyticus Minegishi &al., 2015
Halococcus dombrowski Stan-Lotter &al., 2002
Halococcus hamelinii Goh &al., 2006
Halococcus morrhuae (Farlow, 1880) Kocur & Hodgkiss, 1973
Halococcus qingdaogense Wang &al., 1990
Halococcus saccharolyticus Montero &al., 1990
Halococcus salifodinae Denner &al.,, 1994
Halococcus salsus Chen &al., 2018
Halococcus sediminicola Yim &al., 2014
Halococcus thailandensis Namwong &al., 2007
Haloferax alexandrinus Asker & Ohta, 2002
Haloferax chudinovii Saralov &al., 2014
Haloferax dentrificans (Tomlinson &al., 1986) Tindall &al., 1989
Haloferax elongans Allen &al., 2008
Haloferax gibbonsii Juez &al., 1986
Haloferax larsenii Xu &al., 2007
Haloferax lucentense (Rodriguez-Valera &al., 1983) Torreblanca &al., 1987
Haloferax mediterranei Asker &Ohta, 2002
Haloferax mucosum Allen &al., 2008
Haloferax namakaokahaiae McDuff &al., 2017
Haloferax prahovense Enache &al., 2007
Haloferax profundi Zhang &al., 2020
Haloferax sulfurifontis Elshahed &al., 2004
Haloferax volcanii (Mullakhanbhai & Larsen, 1975) Torreblanca &al., 1986
Halogeometricum borinquense ?
Halogranum amylolyticum Cui &al., 2011
Halogranum gelatinilyticum Cui &al., 2011
Halogranum rubrum Cui &al., 2010
Halogranum salarium Kim &al., 2012
Halolamina litorea Xu &al., 2016
Halolamina pelagica Cui &al., 2011
Halolamina rubra Chu &al., 2014
Halolamina salifodinae Zhang &al., 2013
Halolamina salina Zhang &al., 2013
Halolamina sediminis Koh &al., 2015
Halomarina oriensis Inoue &al., 2011
Halomarina rubra Zhou &al., 2018
Halomarina salina Xu &al., 2016
Halomicrobium katesii Kharroub &al., 2008
Halomicrobium mukohataei (Ihara &al., 1997) Oren &al., 2002
Halomicrobium zhouii Yang & Cui, 2012
Halonotius aquaticus Duran-Viseras &al., 2019
Halonotius pteroides Burns &al., 2010
Halonotius roseus Duran-Viseras &al., 2020
Halonotius terrestris Duran-Viseras &al., 2020
Halopelagius fulvigenes Liu &al., 2013
Halopelagius inordinatus Cui &al., 2010
Halopelagius longus Zhang &al., 2013
Halopiger aswanensis Hezayen &al., 2010
Halopiger djelfimassiliensis Hassani &al., 2016
Halopiger goleimassiliensis Ikram &al., 2016
Halopiger salifodinae Zhang &al., 2013
Halopiger thermotolerans Minegishi &al., 2016
Halopiger xanaduensis Gutierrez &al., 2017
Haloplanus aerogenes Cui &al., 2011
Haloplanus litoreus Han & Cui, 2014
Haloplanus natans Bardavid &al., 2007
Haloplanus rallus Cho &al., 2018
Haloplanus ruber Han & Cui, 2014
Haloplanus rubicundus Kim &al., 2020
Haloplanus salinarum Hwang &al., 2007
Haloplanus salinus Qiu &al., 2014
Haloplanus vescus Cui &al., 2010
Haloquadratum walsbyi Burns &al. 2007
Candidatus Haloredivivus sp. Baker &al., 2006; Golyshina &al., 2017a
Halorhabdus rudnickae Albuquerque &al. 2016
Halorhabdus tiamatea Antunes &al. 2008
Halorhabdus tiamatea Waino &al. 2000
Halorientalis brevis Yuan &al., 2015
Halorientalis pallida Duran-Viseras &al., 2019
Halorientalis persicus Amoozegar &al., 2014
Halorientalis regularis Cui &al., 2011
Halorubrum aethiopicum Cui &al., 2018
Halorubrum aidingense Cui &al., 2016
Halorubrum alkaliphilum Feng &al., 2005
Halorubrum amylolyticum Sun &al., 2020
Halorubrum aquaticum Gutierrez &al., 2011
Halorubrum californiense Presenti &al., 2008
Halorubrum cibi Roh & Bae, 2009
Halorubrum coriense (Kamekura Dyall-Smith, 1996) Oren & Ventosa, 1996
Halorubrum depositum Cehn &al., 2019
Halorubrum distributum (Zvyagintseva & Tarasov 1989) Oren & Ventosa, 1996
Halorubrum ejinorense Castillo &al., 2007
Halorubrum ezzemoulense Kharroub &al., 2006
Halorubrum gandharaense Kondo &al., 2015
Halorubrum glutamatedens Xu &al., 2019
Halorubrum halodurans Corral &al., 2016
Halorubrum kocurii Gutierrez &al., 2008
Halorubrum lacusprofundi (Franzmann &al., 1989) McGenity & Grant 1996
Halorubrum laminarie Han & Cui, 2015
Halorubrum lipolyticum Cui &al., 2006
Halorubrum luteum Hu &al., 2008
Halorubrum orientale Castillo &al., 2006
Halorubrum pallidum Chen &al., 2016
Halorubrum persicum Corral &al., 2015
Halorubrum rubrum Qiu &al., 2014
Halorubrum rutilum Yin &al., 2016
Halorubrum saccharovorum (Tomlinson & Hochstein, 1977), McGenity & Grant, 1996
Halorubrum salinum Zhang & Cui, 2014
Halorubrum salsamenti Chen &al., 2019
Halorubrum sodomense (Oren, 1983) McGenity & Grant, 1996
Halorubrum tebenquichense Lizama &al., 2002
Halorubrum terrestre Ventosa &al., 2004
Halorubrum tibetense Fan &al., 2004
Halorubrum trapanicum (Petter, 1931) McGenity & Grant, 1996
Halorubrum trueperi Chen &al., 2017
††‡ Halorubrum vacuolatum (Mwatha & Grant, 1993) Kamekura &al., 1997; Oren, 2002
Halorubrum xinjiangense Fen &al., 2016
Halorubrum yunnanense Chen &al., 2015
Halorussus amylolyticus Yuan &al., 2015
Halorussus halophilus Ding &al., 2020
Halorussus litoreus Han &al., 2019
Halorussus rarus Cui &al., 2014
Halorussus ruber Xu &al., 2015
Halorussus salinus Xu &al., 2017
Halosimplex carlsbadense Vreeland &al., 2003
Halosimplex litoreum Yuan &al., 2015
Halosimplex pelagicum Han & Cui, 2014
Halosimplex rubrum Han & Cui, 2014
Halostagnicola alkaliphila Nagaoka &al., 2011
Halostagnicola bangensis Corral &al., 2015
Halostagnicola kamekurae Nagaoka &al., 2010
Halostagnicola larsenii Castillo &al., 2006
Haloterrigena daquingensis Wang &al., 2010
Haloterrigena hispanica Romano &al., 2007
Haloterrigena jeotgali Roh &al., 2009
Haloterrigena limicola Cui &al., 2006
Haloterrigena long Cui &al., 2006
Haloterrigena mahii Ding &al., 2017
Haloterrigena saccharevitans Xu &al., 2005
Haloterrigena salifodinae Chen &al., 2020
Haloterrigena salina Gutierrez &al., 2008
Haloterrigena thermotolerans Montalvo-Rodriguez &al., 2000
Haloterrigena turkmenica (Zvyagintseva & Tarasov, 1989) Ventosa &al., 1999
Halovivax asiaticus Castillo &al., 2006
Halovivax cerinus Amoozegar &al., 2015
Halovivax limisalsi Amoozegar &al., 2014
Halovivax ruber Castillo &al., 2007
Helicobacter pylori
*† Hydrogenobacter hydrogenophilus (Kryukov &al., 1984) Stohr &al., 2001
*† Hydrogenobacter thermophilus Kawasumi &al., 1984
*† Hydrogenobaculum acidophilum (≤70°C) (Hydrogenobacter acidophilus) Fashola &al., 2015
Iainarchaeum andersonii Rinke, &al., 2003; Baker &al., 2006
Leptospirillum ferriphilum Lehtovirta-Morley &al., 2011 & 2014; Zhang &al., 2016
Leptospirillum ferro(o)xidans Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
Leptospirillum thermoferrooxidans Johnson, 1998
Candidatus Mancarchaeum acidiphilum Baker &al., 2006; Golyshina &al., 2017a
*† Metallosphaera cuprina Liu &al., 2011; Zhang &al., 2016
*† Metallosphaera hakonensis (Takayanagi &al., 1996) Kurosawa &al., 2003; Zhang &al., 2016
*† Metallosphaera prunae Fuchs &al., 1996; Johnson, 1998; Zhang &al., 2016
*† Metallosphaera sedula Huber &al., 1989; Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
*† Metallosphaera tengchongensis Peng &al., 2015
*† Metallosphaera yellowstonensis Kozubal &al., 2011; Zhang &al., 2016
* Methanocaldocococcus jannaschii
(Methanococcus jannaschii)
Brock &al., 1972
* Methanocaldocococcus thermolithotrophicus Brock &al., 1972
* Methanopyrus kandleri Kurr &al., 1992
Candidatus Micrarchaeum acidiphilum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017a
*† Nanoarchaeum equitans Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017a
Nanobsidianus stetteri
Candidatus Nanopusillus acidilobi Wurch &al., 2016
*†‡ Candidatus Nanosalina sp. Baker &al., 2006; Baker &al., 2010
Candidatus Nanosalinarum sp. Baker &al., 2006; Baker &al., 2010
Natrialba aegyptica Hezayen &al., 2001
Natrialba asiatica
Natrialba chahannaoensis Xu &al., 2001
Natrialba hulunbeirensis Xu &al., 2001
††‡+ Natrialba magadii (Tindall &al., 1984) Kamekura &al., 1997
Natrialba swarupiae Kajale &al., 2020
Natrialba taiwanensis/i> Hezayen &al., 2001; Oren, 2002
Natrinema altunense Xu &al., 2005
Natrinema ejinorense Castillo &al., 2006
Natrinema gari Tapingkae &al., 2008
Natrinema pallidum XMcGenity &al., 1998
Natrinema salaciae Albuquerque &al., 2012
Natrinema soli Rasooli &al., 2017
Natrinema versiforme Xin &al., 2000
Natrarchaeobaculum aegyptiacum (Zhao &al., 2018) Sorokin &al., 2020
Natronoarchaeum sulfireducens (Sorokin &al., 2019) Sorokin &al., 2020
Natronoarchaeum persicum Naghoni &al., 2017
Natronoarchaeum philippinense Shimane &al., 2013
Natronoarchaeum rubrumm Qiu &al., 2014
††‡ Natronobacterium gregoryi Tindall &al. 1984; Oren, 2002
Natronobacterium texcoconense Ruiz-Romero &al. 2013
Natronococcus amylolyticus Kanai &al. 1995
Natronococcus jeotgali Roh &al. 2007
††‡ Natronococcus occultus Tindall &al. 1984
Natronococcus roseus Corral &al. 2013
Natrolimnohabitans innermongolicus Itoh &al. 2005
Natrolimnobius baerhuensis Itoh &al. 2005
Natronomonas gomsonensis Kim &al. 2014
Natronomonas halophila Yin &al. 2020
Natronomonas moolapensis Burns &al. 2010
††‡ Natronomonas pharaonis (Soliman & Truper, 1983) Kamekura &al. 1997
Natronomonas salina Yin &al. 2020
Natronomonas salsuginis Duran-Viseras &al. 2020
Natronorubrum aibiense Cui &al. 2006
Natronorubrum bangense Xu &al. 1999
Natronorubrum halophilum Tao &al. 2020
Natronorubrum sediminis Gutierrez &al. 2010
Natronorubrum sulfidifaciens Cui &al. 2007
Natronorubrum texcoconense Ruiz-Romero &al. 2013
Natronorubrum tibetense Xu &al. 1999
Candidatus Nitrosoarchaeum koreensis Herbold &al., 2017
Candidatus Nitrosoarchaeum limnia Herbold &al., 2017
Candidatus Nitrosocosmicus oleophilus Herbold &al., 2017
Candidatus Nitrosopelagicus brevis Herbold &al., 2017
Candidatus Nitrosopumilus maritimus Lehtovirta-Morley &al., 2011 & 2014; Herbold &al., 2017
Candidatus Nitrosopumilus viennensis Lehtovirta-Morley &al., 2011 & 2014
Candidatus Nitrosopumilus sp. SJ Herbold &al., 2017
Candidatus Nitrosopumilus sp. NF5 Herbold &al., 2017
Candidatus Nitrosopumilus sp. AR1 Herbold &al., 2017
Candidatus Nitrosopumilus sp. AR2 Herbold &al., 2017
Candidatus Nitrosopumilus salaria Herbold &al., 2017
Candidatus Nitrosopumilus sp. D3C Herbold &al., 2017
Candidatus Nitrososphaera evergladensis Herbold &al., 2017
Candidatus Nitrososphaera viennensis Herbold &al., 2017
Candidatus Nitrososphaera gargensis Herbold &al., 2017
Candidatus Nitrosotalea okcheonensis Herbold &al., 2017
Candidatus Nitrosotalea sinensis Herbold &al., 2017
Candidatus Nitrosotalea devanaterra Lehtovirta-Morley &al., 2011 & 2014; Herbold &al., 2017
Candidatus Nitrosotalea bavarica Herbold &al., 2017
Candidatus Nitrosotenuis uzonensis Herbold &al., 2017
Candidatus Nitrosotenuis chungbukensis Herbold &al., 2017
Candidatus Nitrosotenuis cloacae Herbold &al., 2017
Candidatus Parvarchaeum acidiphilum Baker &al., 2006; Baker &al., 2010
Candidatus Parvarchaeum acidophilus Baker &al., 2006; Baker &al., 2010
Picrophilus sp. (Baker-Austin & Dopson, 2007)
Picrophilus oshimae Rampelotto, 2013, Johnson, 1998; Zhang &al., 2016
Picrophilus torridus Rampelotto, 2013, Johnson, 1998; Zhang &al., 2016
* Pyrococcus abyssi Erauso &al., 1993
* Pyrococcus endeavori ?
* Pyrococcus furiosus Erauso &al., 1993
* Pyrococcus glycovorans ?
* Pyrococcus horikoshii Erauso &al., 1993
* Pyrococcus woesei ?
* Pyrolobus fumarii Bloch &al., 1997
Salarchaeum japonicum Shimane &al. 2011
*† Stygiolobus azoricus Segerer &al., 1991; Johnson, 1998; Zhang &al., 2016
*†° Sulfobacillus acidophilus Norris &al., 1996; Johnson, 1998; Fashola &al., 2015
*†° Sulfobacillus benefaciens Johnson &al., 2009
*†° Sulfobacillus sibiricus Melamud &al., 1996
*†° Sulfobacillus thermosulfidooxidans Golovacheva & Karavaiko, 1991
*†° Sulfobacillus thermotolerans Bogdanova &al., 2006
*† Sulfolobus acidocaldarius Brock &al., 1972; Johnson, 1998; Edwards &al., 2000; Zhang &al., 2016
Sulfolobus hakonensis Johnson, 1998
*† Sulfolobus islandicus Brock &al., 1972
* Sulfolobus metallicus Brock &al., 1972; Johnson, 1998; Fashola &al., 2015; Zhang &al., 2016
Sulfolobus mirabilis Johnson, 1998
*† Sulfolobus neozealandicus Brock &al., 1972
Sulfolobus rivotinct Zhang &al., 2016
*† Sulfolobus shibatae Brock &al., 1972; Johnson, 1998; Zhang &al., 2016
*† Sulfolobus solfataricus Brock &al., 1972; Johnson, 1998, Dopson (2003); Zhang &al., 2016
*† Sulfolobus tengchongensis Brock &al., 1972; Zhang &al., 2016
*† Sulfolobus thuringiensis Brock &al., 1972
*† Sulfolobus tokodaii Brock &al., 1972; Dopson 2003; Zhang &al., 2016
*† Sulfolobus yangmingensis Brock &al., 1972; Zhang &al., 2016
* Sulfolobus sp. AMP12/99 ?
* Sulfolobus sp. CH7/99 ?
* Sulfolobus sp. FF5/00 ?
* Sulfolobus sp. G81 ?
* Sulfolobus sp. H ?
* Sulfolobus sp. JP2 ?
* Sulfolobus sp. JP3 ?
* Sulfolobus sp. K00 8-41 ?
* Sulfolobus sp. L00 11 ?
* Sulfolobus sp. L00 24 ?
* Sulfolobus sp. M02 ?
* Sulfolobus sp. MV2/99 ?
* Sulfolobus sp. MVSoil3/SC2 ?
* Sulfolobus sp. MVSoil6/SC1 ?
* Sulfolobus sp. NGB6/00 ?
* Sulfolobus sp. NGB23/00 ?
* Sulfolobus sp. NGL8/00 ?
* Sulfolobus sp. NO82 ?
* Sulfolobus sp. NOB8H2 ?
* Sulfolobus sp. RC3 ?
* Sulfolobus sp. RC6/00 ?
* Sulfolobus sp. RCSC1/01 ?
* Sulfolobus sp. Sko-3 ?
* Sulfolobus sp. ?
* Sulfolobus sp. Ta ?
* Sulfolobus sp. Tu A ?
* Sulfolobus sp. Tu B-1 ?
* Sulfolobus sp. Y00 58-82 ?
* Sulfolobus sp. Y01 90#-18 ?
* Sulfolobus sp. Y99 9-16 ?
* Sulfolobus sp. Y99 9-23 ?
* Sulfolobus sp. (uncultured) ?
Sulfurisphaera ohwakuensis Zhang &al., 2016
Sulfurococcus mirabilis Zhang &al., 2016
Sulfurococcus yellowstonensis Zhang &al., 2016
Sulfurococcus yellowstonii Johnson, 1998
Telmatobacter bradus ()
* Thermococcus acidaminovorans ?
* Thermococcus aegaeus ?
* Thermococcus aggregans ?
*†† Thermococcus alcaliphilus Keller &al., 1997
* Thermococcus atlanticus ?
*** Thermococcus barophilus Marteinsson &al., 1999
* Thermococcus barossi ?
*† Thermococcus celer Zillig, 1983
*** Thermococcus chitonophagus
(one of only three archaeal chiton-philes)
Huber & Stetter, 1996
* Thermococcus coalescens ?
* Thermococcus fumicolans ?
* Thermococcus gammatolerans Jolivet, 2003
* Thermococcus gorgonarius ?
* Thermococcus guaymasensis ?
*** Thermococcus hydrothermalis Godfroy &al., 1997
* Thermococcus kodakarensis Atomi &al., 2004
***† Thermococcus litoralis Neuner &al., 2001
* Thermococcus marinus ?
* Thermococcus mexicalis ?
* Thermococcus pacificus ?
*** Thermococcus peptonophilus (more thermophilic at higher pressures and more barophilic at higher temps Canganella &al., 1996
* Thermococcus profundis Kobayashi & Horikoshi, 1995
* Thermococcus radiotolerans ?
* Thermococcus sibiricus ?
* Thermococcus siculi ?
* Thermococcus stetteri Miroshnichenko, 1990
* Thermococcus waimanguensis ?
* Thermococcus waiotapuensis ?
*††? Thermococcus zilligii Zillig, 1982
* Thermococcus sp. ?
* Thermococcus sp. 12-4 NCBI’s Taxonomy Browser
* Thermococcus sp. 13-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 13-3 NCBI’s Taxonomy Browser
* Thermococcus sp. 21-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 23-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 23-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 26-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 26-3 NCBI’s Taxonomy Browser
* Thermococcus sp. 28-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 29-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 300-Tc NCBI’s Taxonomy Browser
* Thermococcus sp. 30-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 31-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 31-3 NCBI’s Taxonomy Browser
* Thermococcus sp. 5-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 70-4-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 83-5-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 9N2 NCBI’s Taxonomy Browser
* Thermococcus sp. 9N3 NCBI’s Taxonomy Browser
* Thermococcus sp. 9oN-7 NCBI’s Taxonomy Browser
* Thermococcus sp. AEPII 1a NCBI’s Taxonomy Browser
* Thermococcus sp. AM4 NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete70-I78 NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete70-SCI NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete85-I78 NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete 85-SCI NCBI’s Taxonomy Browser
* Thermococcus sp. AT1273 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-17 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-27 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-39 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-45 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-2 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-37 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-39 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-3 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-61 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-62 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-65 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax98-43 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax98-46 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax98-48 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax99-47 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax99-57 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax99-67 NCBI’s Taxonomy Browser
* Thermococcus sp. B1001 NCBI’s Taxonomy Browser
* Thermococcus sp. BHI60a21 NCBI’s Taxonomy Browser
* Thermococcus sp. BHI80a28 NCBI’s Taxonomy Browser
* Thermococcus sp. BHI80a40 NCBI’s Taxonomy Browser
* Thermococcus sp. CAR-80 NCBI’s Taxonomy Browser
* Thermococcus sp. CKU-199 NCBI’s Taxonomy Browser
* Thermococcus sp. CL1 NCBI’s Taxonomy Browser
* Thermococcus sp. CL2 NCBI’s Taxonomy Browser
* Thermococcus sp. CMI NCBI’s Taxonomy Browser
* Thermococcus sp. CNR-5 NCBI’s Taxonomy Browser
* Thermococcus sp. CX1 NCBI’s Taxonomy Browser
* Thermococcus sp. CX2 NCBI’s Taxonomy Browser
* Thermococcus sp. CX3 NCBI’s Taxonomy Browser
* Thermococcus sp. CX4 NCBI’s Taxonomy Browser
* Thermococcus sp. CYA NCBI’s Taxonomy Browser
* Thermococcus sp. Dex80a71 NCBI’s Taxonomy Browser
* Thermococcus sp. Dex80a75 NCBI’s Taxonomy Browser
* Thermococcus sp. enrichment clone SA3 NCBI’s Taxonomy Browser
* Thermococcus sp. GE8 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda2 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda3 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda4 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda5 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda6 NCBI’s Taxonomy Browser
* Thermococcus sp. GT NCBI’s Taxonomy Browser
* Thermococcus sp. GU5L5 NCBI’s Taxonomy Browser
* Thermococcus sp. JDF-3 NCBI’s Taxonomy Browser
* Thermococcus sp. KI NCBI’s Taxonomy Browser
* Thermococcus sp. KS-1 NCBI’s Taxonomy Browser
* Thermococcus sp. KS-8 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1031 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1049 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1083 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1092 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1099 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ10 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ11 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ12 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ13 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ1 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ2 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ3 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ5 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ6 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ8 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ9 NCBI’s Taxonomy Browser
* Thermococcus sp. NA1 NCBI’s Taxonomy Browser
* Thermococcus sp. NS85-T NCBI’s Taxonomy Browser
* Thermococcus sp. OGL-20P NCBI’s Taxonomy Browser
* Thermococcus sp. P6 NCBI’s Taxonomy Browser
* Thermococcus sp. Pd70 NCBI’s Taxonomy Browser
* Thermococcus sp. Pd85 NCBI’s Taxonomy Browser
* Thermococcus sp. Rt3 NCBI’s Taxonomy Browser
* Thermococcus sp. SB611 NCBI’s Taxonomy Browser
* Thermococcus sp. SN531 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-1-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-1-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-1-95 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-2-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-2-95 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-365-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-365-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-365-95 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-4-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-4-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-CRC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-CRC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-MC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-SC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-SC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-0 age SC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-CRC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-CRC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-MC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-MC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-SC-ISCS NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-SC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-SC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-CRC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-CRC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-MC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-MC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-SC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-I-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-I-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-S-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-S-85 NCBI’s Taxonomy Browser
* Thermococcus sp. TK1 NCBI’s Taxonomy Browser
* Thermococcus sp. TM1 NCBI’s Taxonomy Browser
* Thermococcus sp. TS2 NCBI’s Taxonomy Browser
* Thermococcus sp. TS3 NCBI’s Taxonomy Browser
* Thermococcus sp. vp197 NCBI’s Taxonomy Browser
* Thermococcus sp. (uncultured) ?
Thermogymnomonas acidocola ?
Thermoplasma acidophilum Johnson, 1998; Zhang &al., 2016
Thermoplasma volcanium Johnson, 1998; Zhang &al., 2016
* Thermotoga caldifontis ?
* Thermotoga elfii ?
* Thermotoga hypogea ?
* Thermotoga lettingae ?
* Thermotoga maritima ?
* Thermotoga napthophila ?
* Thermotoga neapolitana ?
* Thermotoga petrophila ?
* Thermotoga profunda ?
* Thermotoga subterranea ?
* Thermotoga thermarum ?
Thiobacillus acidophilus ()
Thiobacillus dentrificans (Fashola &al., 2015)
Thiobacillus organovorus ()
Thiobacillus prosperus Johnson, 1998
Thiomonas cuprina
(Thiobacillus cuprinus)
Johnson, 1998



References

Note: This section is still under construction!


Schuerger & Nicholson, 2016.
http://doi.org/10.1089/ast.2016.1587

Nicholson &al., 2013.
https://doi.org/10.1073/pnas.1209793110

Baker B.J., Tyson G.W., Webb R.I., Flanagan J., Hugenholtz P., Allen E.E., Banfield J.F., (2006); Lineages of acidophilic archaea revealed bycommunity genomic analysis. Science. 2006 Dec 22;314(5807):1933-5. doi: 10.1126/science.1132690 <https://pubmed.ncbi.nlm.nih.gov/17185602/>

Edwards K.J., Bond P.L., Gihring T.M., Banfield J.F., (2000); An Archaeal Iron-Oxidizing Extreme Acidophile Important in Acid Mine Drainage. 10 MARCH 2000 VOL 287 SCIENCE p. 1796-1799. <http://science.sciencemag.org/content/287/5459/1796.full>

Fashola M.O., Babalola O.O., Ngole V.M., (2015); Diversity of Acidophilic Bacteria and Archaea and their Roles in Bioremediation of Acid Mine Drainage. January 2015. British Microbiology Research Journal 8(3):443-456. DOI: 10.9734/BMRJ/2015/14365. <https://www.researchgate.net/publication/276456404_...>

Golyshina O.V., Kublanov I.V., Tran H., Korzhenkov A.A., Lünsdorf H., Nechitaylo T.Y., Gavrilov S.N., Toshchakov S.V., Golyshin P.N., (2016); Biology of archaea from a novel family Cuniculiplasmataceae (Thermoplasmata) ubiquitous in hyperacidic environments. Scientific Reports 6, Article number: 39034 (2016) Received: 07 September 2016, Accepted: 16 November 2016, Published online: 14 December 2016 doi:10.1038/srep39034 <https://www.nature.com/articles/srep39034>

Golyshina O.V., Toshchakov S.V., Makarova K.S., Gavrilov S.N., Korzhenkov A.A., LaCono V., Arcadi E., Nechitaylo T.Y., Ferrer M., Kublanov I.V., Wolf Y.I., Yakimov M.M., Golyshin, P.N., (2017) 'ARMAN' archaea depend on association with euryarchaeal host in culture and in situ. 2017. DOI: 10.1038/s41467-017-00104-7 <https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5498576/>

Golyshina O.V., Tran H., Reva O.N., Lemak S., Yakunin A.F., Goesmann A., Nechitaylo T.Y., LaCono V., Smedile F., Slesarev A., Rojo D., Barbas C., Ferrer M., Yakimov M.M., Golyshin P.N., (2017); Metabolic and evolutionary patterns in the extremely acidophilic archaeon Ferroplasma acidiphilum YT. Scientific Reports volume 7, Article number: 3682 <https://www.nature.com/articles/s41598-017-03904-5>

He Z.-G., Zhong H., Li Y., (2004); Acidianus tengchongensis sp. nov., a new species of acidothermophilic archaeon isolated from an acidothermal spring. Curr Microbiol. 2004 Feb;48(2):159-63. doi: 10.1007/s00284-003-4155-9. <https://pubmed.ncbi.nlm.nih.gov/15057486/>

Herbold C.W., Lehtovirta-Morley L.E., Jung M.-Y., Jehmlich N., Hausmann B., Ping Han, Loy A., Pester P., Sayavedra-Soto L.A., Rhee S.-K., Prosser J.I., Nicol H.W., Wagner M., Gubry-Rangin C., (2017); Ammonia-oxidising archaea living at low pH: Insights from comparative genomics. Environmental Microbiology (2017) 19(12), 4939–4952. Environment Microbiology. Volume 19, Issue 12. Thematic Issue: Microbial N2O Sources and Sinks. December 2017. Pages 4939-4952. <https://doi.org/10.1111/1462-2920.13971>

Johnson D.B., (1998); Biodiversity and ecology of acidophilic microorganisms. FEMS Microbiology Ecology, Volume 27, Issue 4, December 1998, Pages 307-317. <https://doi.org/10.1111/j.1574-6941.1998.tb00547.x>

Lehtovirta-Morley L.E., Stoecker K., Vilcinskas A., Prosser J.I., Nicol G.W., (2011); Cultivation of an obligate acidophilic ammonia oxidizer from a nitrifying acid soil. PNAS vol. 108 no. 38, 15892–15897, doi: 10.1073/pnas.1107196108. <https://doi.org/10.1073/pnas.1107196108>

Lehtovirta-Morley L.E., Ge C., Ross J., Yao H., Nicol G.W., Prosser J.I., (2014);

Oren A., (2008); Microbial life at high salt concentrations: phylogenetic and metabolic diversity. Saline Syst. (2008); 4:2. Published online 2008 Apr 15. doi: 10.1186/1746-1448-4-2 <https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2329653/>

Plumb J.J., Haddad C.M., Gibson J.A., Franzmann P.D., (2007); Acidianus sulfidivorans sp. nov., an extremely acidophilic, thermophilic archaeon isolated from a solfatara on Lihir Island, Papua New Guinea, and emendation of the genus description. Int J Syst Evol Microbiol. 2007 Jul;57(Pt 7):1418-1423. doi: 10.1099/ijs.0.64846-0. <https://pubmed.ncbi.nlm.nih.gov/17625168/>

Zhang B., Wu Y.-F., Song J.-L., Huang Z.-S., Liu S.-J., Jiang C.-Y., (2015); Alicyclobacillus fodiniaquatilis sp. nov., isolated from acid mine water. Int J Syst Evol Microbiol. 2015 Dec;65(12):4915-4920. doi: 10.1099/ijsem.0.000695. <https://pubmed.ncbi.nlm.nih.gov/26476812/>.

Zhang W.-Y., Hu J., Zhang X.-Q., Zhu X.-F., Wu M., (2015); Bacillus salitolerans sp. nov., a novel bacterium isolated from a salt mine in Xinjiang province, China. Antonie Van Leeuwenhoek. 2015 Aug;108(2):443-51. doi: 10.1007/s10482-015-0497-0. Epub 2015 Jun 16. <https://pubmed.ncbi.nlm.nih.gov/26076748/>

Zhang L., (2007); Bacillus decisifrondis sp. nov., isolated from soil underlying decaying leaf foliage. Int J Syst Evol Microbiol. 2007 May;57(Pt 5):974-978. doi: 10.1099/ijs.0.64440-0. <https://pubmed.ncbi.nlm.nih.gov/17473244/>.

Zhang R., (2016); Biofilm formation and extracellular polymeric substances of acidophilic metal/sulfur-oxidizing archaea. Duisburg-Essen Publications online <https://duepublico2.uni-due.de/receive/duepublico_mods_00040230>.

Zhang Y.-G., (2018); Bacillus tamaricis sp. nov., an alkaliphilic bacterium isolated from a Tamarix cone soil. Int J Syst Evol Microbiol. 2018 Feb;68(2):558-563. doi: 10.1099/ijsem.0.002543. Epub 2018 Jan 3. <https://pubmed.ncbi.nlm.nih.gov/29297846/>.



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