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Euryarchaeota



Introduction

This page is under construction.




Definitons

Acidophiles: This group includes organisms capable of thriving in hyperacidic conditions, most of which have an optimal growth pH of 5 or lower (OGpH<5)

Hyperacidotolerant, near-acidophilic neutriphiles: This group includes organisms that have an OGpH slightly above this (e.g., Nanoarchaeum equitans, OGpH=6) but can nonetheless be found flourishing in hyperacidic environments (pH=0.5-1.5).




Taxonomic/Phylogenetic Overview of Hypergeophilic & Hypergeotolerant Genera

“Hypergeophilic genera” and “hypergeotolerant genera” refer here to genera known to include hypergeophilic species and/or species with hypergeotolerances.

Genera containing thermophilic and hyperthermotolerant organisms are marked with an asterisk (*)

Genera containing barophilic and hyperbarotolerant organisms are marked with two asterisks (**)

Genera containing acidophilic and hyperacidotolerant organisms are marked with a dagger (†)

Genera containing alkaliphilic and hyperalkalitolerant organisms are marked with two daggers (††)

Genera containing thermoacidophilic and hyperthermoacidotolerant organisms are marked with an asterisk and a dagger (*†)

Genera containing halophilic and hyperhalotolerant organisms are marked with a double dagger (‡)

Bacteria marked with an asterisk and a degree sign (*°) are endospore-forming polyextremotolerant organisms. Bacterial endospores tend to give the bacteria capable of producing them polyextremotolerance. Endospores are known for being able to survive extremely high doses of radiation, vacuum dessication, extreme heat and cold, lack of nutrients, and in some cases even the passage of hundreds of millions of years (for example, Virgibacillus marismortui (formerly Bacillus marismortui and Bacillus strain 2-9-3) was revived by scientists from 250-million-year-old spores (Vreeland, 2000); for comparison, the first dinosaurs began appearing about 220 MYA — 30 million years later). Although these endospore-forming bacteria can survive a variety of harsh conditions, endospore-formation is not done for reproduction, as with fungal spore formation, but is instead a last-ditch effort to survive conditions the bacterium would not otherwise be able to, which means these critters aren’t true polyextremophiles, because they don’t (necessarily) actually grow better in extreme conditions, and in their endospore form don’t even grow at all.

In addition to other sources cited, much of this information was compiled with the help of NCBI’s Taxonomy Browser.


Domain: Archaea
Phylum: Euryarchaeota
Class: Archaeoglobi
Order: Archaeoglobales
Family: Archaeoglobaceae
Genus: Archaeoglobus *
Class: Stenosarchaea
Order: Candidatus Nanohaloarchaeota
Family: Nanohaloarchaea
Genus: Haloredivivus †‡
Genus: Nanosalina *†‡
Genus: Nanosalinarum †‡
Class: Halobacteria
Order: Halobacteriales
Family: Haloarculaceae
Genus: Haloarcula
Genus: Halorhabdus
Genus: Halorientalis
Genus: Halosimplex
Genus: Natronomonas ††‡
Family: Halobacteriaceae
Genus: Haladaptatus
Genus: Halalkalicoccus ††‡
Genus: Halarchaeum †‡
Genus: Haloalcalophilum ††‡
Genus: Halobacterium
Genus: Halomarina
Genus: Halorussus
Genus: Natronoarchaeum
Genus: Salarchaeum
Family: Halococceae
Genus: Halococcus
Order: Haloferacales
Family: Haloferacaceae
Genus: Halobellus
Genus: Haloferax
Genus: Halopelagius
Genus: Haloplanus
Genus: Haloquadratum
Family: Halorubraceae
Genus: Halobaculum
Genus: Halogeometricum
Genus: Halogranum
Genus: Halolamina
Genus: Halonotius
Genus: Halorubrum ††‡
Order: Natrialbales
Family: Natrialbaceae
Genus: Halobiforma
Genus: Halopiger
Genus: Halostagnicola
Genus: Haloterrigena
Genus: Halovivax
Genus: Natrarchaeobaculum
Genus: Natrialba ††‡+
Genus: Natrinema
Genus: Natronobacterium ††‡
Genus: Natronococcus ††‡
Genus: Natronolimnohabitans
Genus: Natronolimnobius
Genus: Natronorubrum
Class: Methanococci
Order: Methacoccales
Family: Methanococcaceae
Genus: Methanocaldococcus ***
Class: Methanopyri
Order: Methanopyrales
Family: Methanopyraceae
Genus: Methanopyrus *
Class: Thermococci
Order: Thermococcales
Family: Thermococcaceae
Genus: Pyrococcus ***
Genus: Thermococcus ***†††
Class: Thermoplasmata
Order: Methanomassiliicoccales
Family: Methanomassiliicoccaceae
Genus: Methanomassiliicoccus
Order: Thermoplasmatales
Family: Cuniculiplasmaceae
Genus: Cuniculiplasma
Family: Ferroplasmaceae
Genus: Acidiplasma
Genus: Ferroplasma
Family: Picrophilaceae
Genus: Picrophilus
Family: Incertae sedis
Genus: Thermogymnomonas
Superphylum: DPANN group (9x; 3*, 9†, 3*†)
Phylum: Candidatus Diapherotrites (1x; 1†)
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Iainarchaeum (1x; 1†)
Phylum: Aenigmarchaeota (1x; 1†)
Class: (unnamed?)
Order: Aenigmarchaeales (1x; 1†)
Family: (unnamed?)
Genus: Aenigmarchaeum (1x; 1†)
Phylum: Nanoarchaeota (3x; 3*, 3†, 3*†)
Class: (unnamed?)
Order: Nanoarchaeales (3x; 3*, 3†, 3*†)
Family: Nanoarchaeaceae (1x; 1*, 1†, 1*†)
Genus: Nanoarchaeum (1x; 1*, 1†, 1*†)
Family: Nanopusillaceae (2x; 2*, 2†, 2*†)
Genus: Nanobsidianus (1x; 1*, 1†, 1*†)
Genus: Nanopusillus (1x; 1*, 1†, 1*†)
Phylum: Incertae sedis
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Mancarchaeum (1x; 1†)
Phylum: Candidatus Micrarchaeota (1x; 1†)
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Micrarchaeum (1x; 1†)
Phylum: Candidatus Parvarchaeota (2x; 2†)
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Parvarchaeum (2x; 2†)
Kingdom: Proteoarchaeota (93x; 60*, 57†, 24*†)
Superphylum: TACK group (93x; 60*, 57†, 24*†)
Phylum: Thaumarchaeota (23x; 23†)
Class: Nitrososphaeria (15x; 15†)
Order: Nitrosopumilales (11x; 11†)
Family: Nitrosopumilaceae (11x; 11†)
Genus: Candidatus Nitrosoarchaeum (3x; 3†)
Genus: Nitrosopumilus (8x; 8†)
Order: Nitrososphaerales (4x; 4†)
Family: Nitrososphaeraceae (4x; 4†)
Genus: Candidatus Nitrosocosmicus (1x; 1†)
Genus: Candidatus Nitrososphaera (3x; 3†)
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Nitrosopelagicus (1x; 1†)
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Nitrosotalea (4x; 4†)
Class: Incertae sedis
Order: Incertae sedis
Family: Incertae sedis
Genus: Candidatus Nitrosotenuis (3x; 3†)
Phylum: Crenarchaeota (70x; 60*, 34†, 24*†)
Class: Thermoprotei (69x; 60*, 33†, 24*†)
Order: Desulfurococcales (4x; 4*)
Family: Desulfurococcacae (2x; 2*)
Genus: Aeropyrum (2x; 2*)
Family: Pyrodictiacae (2x; 2*)
Genus: Geogemma (1x; 1*)
Genus: Pyrolobus (1x; 1*)
Order: Sulfolobales (65x; 56*, 33†, 24*†)
Family: Sulfobaceae (65x; 56*, 33†, 24*†)
Genus: Acidianus (10x; 10*†)
Genus: Metallosphaera (5x; 5†)
Genus: Stygiolobus (1x; 1*†)
Genus: Sulfolobus (45x; 45*, 13†, 13*†)
Genus: Sulfurisphaera (1x; 1†)
Genus: Sulfurococcus (3x; 3†)
Class: Incertae sedis
Order: Cenarchaeales (1x; 1†)
Family: Cenarchaeaceae (1x; 1†)
Genus: Cenarchaeum (1x; 1†)



Alphabetical List

Described Species
Genus & Species Temperature
T°C range
Pressure
kPa range
Proton
pH range
Water
aw range
Salt
%NaCl range
Radiation
Gy range
low_T°C OGT°C high_T°C low_kPa OGkPa high_kPa min._pH OGpH max._pH low_aw OGaw high_aw low_NaCl OGNaCl high_NaCl low_Gy OGGy high_Gy
Acidiplasma_aeolicum 15 45 65 - - - 0 1.4-1.6 4 - - - - - - - - -
Acidiplasma_cupricumulans 22 53.6 63 - - - 0.4 1-1.2 1.8 - - - - - - - - -



In-Line Citations by Genus & species

Described Species
Genus & Species Source
Acidianus ambivalens Johnson, 1998; Zhang &al., 2015; (Zillig and Böck 1987) Fuchs &al. 1996
Acidianus brierleyi Segerer &al. 1986; Johnson, 1998; Edwards &al., 2000; Fashola &al., 2015; Zhang &al., 2015
Acidianus_convivator Haring &al., 2005
Acidianus copahuensis Giaveno &al., 2012; Zhang &al., 2015; Urbieta &al., 2017
Acidianus infernus Segerer &al. 1986; Johnson, 1998; Zhang &al., 2015
Acidianus manzaensis Yoshida &al., 2006; Ding &al. 2011; Zhang &al., 2015
Acidianus pozzuoliensis Haring &al., 2005
Acidianus sulfidivorans Plumb &al. 2007; Zhang &al., 2015
Acidianus tengchongensis He &al. 2004; Zhang &al., 2015
Acidianus sp. DSM 29099 Zhang &al., 2015
Acidianus sp. RZ1 Zhang &al., 2015
Acidiplasma aeolicum Golyshina &al., 2009; Zhang &al., 2015
Acidiplasma cupricumulans (Hawkes &al., 2008) Golyshina &al., 2009; Zhang &al., 2015
Aeropyrum _camini Nakagawa &al., 2004
Aeropyrum pernix Sako &al., 1996
Archaeoglobus fulgidus Stetter &al., 1988; Klenk &al., 1997
Archaeoglobus infectus Mori &al., 2008
Archaeoglobus lithotrophicus ?
Archaeoglobus profundis Burggraf &al. 1990
Archaeoglobus veneficus Huber &al. 1998
Candidatus Aenigmarchaeum subterraneum Baker &al., 2006; Golyshina &al., 2017
Candidatus Cenarchaeum symbiosum A Herbold &al., 2017
Candidatus Haloredivivus sp. Baker &al., 2006; Golyshina &al., 2017
Candidatus Iainarchaeum andersonii Baker &al., 2006; Golyshina &al., 2017
Candidatus Mancarchaeum acidiphilum Baker &al., 2006; Golyshina &al., 2017
Candidatus Micrarchaeum acidiphilum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Nanoarchaeum equitans Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Nanobsidianus stetteri
Candidatus Nanopusillus acidilobi
Candidatus Nanosalina sp. Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Nanosalinarum sp. Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Nitrosoarchaeum koreensis Herbold &al., 2017
Candidatus Nitrosoarchaeum limnia Herbold &al., 2017
Candidatus Nitrosoarchaeum limnia Herbold &al., 2017
Candidatus Nitrosocosmicus oleophilus Herbold &al., 2017
Candidatus Nitrosopelagicus brevis Herbold &al., 2017
Candidatus Nitrosopumilus maritimus Lehtovirta-Morley &al., 2014; Herbold &al., 2017
Candidatus Nitrosopumilus viennensis Lehtovirta-Morley &al., 2014
Candidatus Nitrosopumilus sp. SJ Herbold &al., 2017
Candidatus Nitrosopumilus sp. NF5 Herbold &al., 2017
Candidatus Nitrosopumilus sp. AR1 Herbold &al., 2017
Candidatus Nitrosopumilus sp. AR2 Herbold &al., 2017
Candidatus Nitrosopumilus salaria Herbold &al., 2017
Candidatus Nitrosopumilus sp. D3C Herbold &al., 2017
Candidatus Nitrososphaera evergladensis Herbold &al., 2017
Candidatus Nitrososphaera viennensis Herbold &al., 2017
Candidatus Nitrososphaera gargensis Herbold &al., 2017
Candidatus Nitrosotalea okcheonensis Herbold &al., 2017
Candidatus Nitrosotalea sinensis Herbold &al., 2017
Candidatus Nitrosotalea devanaterra Lehtovirta-Morley &al., 2011 & 2014; Herbold &al., 2017
Candidatus Nitrosotalea bavarica Herbold &al., 2017
Candidatus Nitrosotenuis uzonensis Herbold &al., 2017
Candidatus Nitrosotenuis chungbukensis Herbold &al., 2017
Candidatus Nitrosotenuis cloacae Herbold &al., 2017
Candidatus Parvarchaeum acidiphilum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Parvarchaeum acidophilus Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Ferroplasma_acidarmanus Edwards &al., 2000, Dopson &al.2003; Zhang &al., 2015
Ferroplasma acidiphilum Fashola &al., 2015; Zhang &al., 2015
Ferroplasma cyprexacervatum ()
Ferroplasma thermophilum Zhang &al., 2015
Geogemma barossi
(Pyrolobus fumarii, strain 121)
Kashefi & Lovley, 2003
Haladaptatus cibarius ()
Haladaptatus litoreus ()
Haladaptatus pallidirubidus ()
Haladaptatus paucihalophilus ()
††‡ Halalkalicoccus jeotgali ()
††‡ Halalkalicoccus tibetensis ()
Halarchaeum acidiphilum ()
Haloalcalophilum atacamensis Oren & Ventosa, 2000
Haloarcula amylolytica ?
Haloarcula argentinensis ?
Haloarcula hispanica ?
Haloarcula japonica ?
Haloarcula marismortui ?
Haloarcula quadrata ?
Haloarcula vallismortis ?
Halobacterium jilantaiense ?
Halobacterium noricense ?
Halobacterium salinarum ?
Halobacterium piscisalsi ?
Halobaculum magnesiipjilum Shimoshige &al., 2013
Halobaculum roseum Chen &al., 2017
Halobaculum saliterrae Myers & King, 2020
Halobellus captivus Chen &al., 2020
Halobellus clavatus Cui &al., 2011
Halobellus inordinatus Qiu &al., 2013
Halobellus Iimi Cui &al., 2012
Halobellus litoreus Zhao &al., 2014
Halobellus ramosii Perez-Davo &al., 2015
Halobellus rarus Zhang &al., 2014
Halobellus refus Cha &al., 2014
Halobellus salinus Cui &al., 2012
Halobiforma haloterrestris ?
Halobiforma lacisalsi ?
Halobiforma nitratireducens ?
Halococcus agarilyticus Minegishi &al., 2015
Halococcus dombrowski Stan-Lotter &al., 2002
Halococcus hamelinii Goh &al., 2006
Halococcus morrhuae (Farlow, 1880) Kocur & Hodgkiss, 1973
Halococcus qingdaogense Wang &al., 1990
Halococcus saccharolyticus Montero &al., 1990
Halococcus salifodinae Denner &al.,, 1994
Halococcus salsus Chen &al., 2018
Halococcus sediminicola Yim &al., 2014
Halococcus thailandensis Namwong &al., 2007
Haloferax alexandrinus Asker & Ohta, 2002
Haloferax chudinovii Saralov &al., 2014
Haloferax dentrificans (Tomlinson &al., 1986) Tindall &al., 1989
Haloferax elongans Allen &al., 2008
Haloferax gibbonsii Juez &al., 1986
Haloferax larsenii Xu &al., 2007
Haloferax lucentense (Rodriguez-Valera &al., 1983) Torreblanca &al., 1987
Haloferax mediterranei Asker &Ohta, 2002
Haloferax mucosum Allen &al., 2008
Haloferax namakaokahaiae McDuff &al., 2017
Haloferax prahovense Enache &al., 2007
Haloferax profundi Zhang &al., 2020
Haloferax sulfurifontis Elshahed &al., 2004
Haloferax volcanii (Mullakhanbhai & Larsen, 1975) Torreblanca &al., 1986
Halogeometricum borinquense ?
Halogranum amylolyticum Cui &al., 2011
Halogranum gelatinilyticum Cui &al., 2011
Halogranum rubrum Cui &al., 2010
Halogranum salarium Kim &al., 2012
Halolamina litorea Xu &al., 2016
Halolamina pelagica Cui &al., 2011
Halolamina rubra Chu &al., 2014
Halolamina salifodinae Zhang &al., 2013
Halolamina salina Zhang &al., 2013
Halolamina sediminis Koh &al., 2015
Halomarina oriensis Inoue &al., 2011
Halomarina rubra Zhou &al., 2018
Halomarina salina Xu &al., 2016
Halomicrobium katesii Kharroub &al., 2008
Halomicrobium mukohataei (Ihara &al., 1997) Oren &al., 2002
Halomicrobium zhouii Yang & Cui, 2012
Halonotius aquaticus Duran-Viseras &al., 2019
Halonotius pteroides Burns &al., 2010
Halonotius roseus Duran-Viseras &al., 2020
Halonotius terrestris Duran-Viseras &al., 2020
Halopelagius fulvigenes Liu &al., 2013
Halopelagius inordinatus Cui &al., 2010
Halopelagius longus Zhang &al., 2013
Halopiger aswanensis Hezayen &al., 2010
Halopiger djelfimassiliensis Hassani &al., 2016
Halopiger goleimassiliensis Ikram &al., 2016
Halopiger salifodinae Zhang &al., 2013
Halopiger thermotolerans Minegishi &al., 2016
Halopiger xanaduensis Gutierrez &al., 2017
Haloplanus aerogenes Cui &al., 2011
Haloplanus litoreus Han & Cui, 2014
Haloplanus natans Bardavid &al., 2007
Haloplanus rallus Cho &al., 2018
Haloplanus ruber Han & Cui, 2014
Haloplanus rubicundus Kim &al., 2020
Haloplanus salinarum Hwang &al., 2007
Haloplanus salinus Qiu &al., 2014
Haloplanus vescus Cui &al., 2010
Haloquadratum walsbyi Burns &al. 2007
Candidatus Haloredivivus sp. Baker &al., 2006; Golyshina &al., 2017
Halorhabdus rudnickae Albuquerque &al. 2016
Halorhabdus tiamatea Antunes &al. 2008
Halorhabdus tiamatea Waino &al. 2000
Halorientalis brevis Yuan &al., 2015
Halorientalis pallida Duran-Viseras &al., 2019
Halorientalis persicus Amoozegar &al., 2014
Halorientalis regularis Cui &al., 2011
Halorubrum aethiopicum Cui &al., 2018
Halorubrum aidingense Cui &al., 2016
Halorubrum alkaliphilum Feng &al., 2005
Halorubrum amylolyticum Sun &al., 2020
Halorubrum aquaticum Gutierrez &al., 2011
Halorubrum californiense Presenti &al., 2008
Halorubrum cibi Roh & Bae, 2009
Halorubrum coriense (Kamekura Dyall-Smith, 1996) Oren & Ventosa, 1996
Halorubrum depositum Cehn &al., 2019
Halorubrum distributum (Zvyagintseva & Tarasov 1989) Oren & Ventosa, 1996
Halorubrum ejinorense Castillo &al., 2007
Halorubrum ezzemoulense Kharroub &al., 2006
Halorubrum gandharaense Kondo &al., 2015
Halorubrum glutamatedens Xu &al., 2019
Halorubrum halodurans Corral &al., 2016
Halorubrum kocurii Gutierrez &al., 2008
Halorubrum lacusprofundi (Franzmann &al., 1989) McGenity & Grant 1996
Halorubrum laminarie Han & Cui, 2015
Halorubrum lipolyticum Cui &al., 2006
Halorubrum luteum Hu &al., 2008
Halorubrum orientale Castillo &al., 2006
Halorubrum pallidum Chen &al., 2016
Halorubrum persicum Corral &al., 2015
Halorubrum rubrum Qiu &al., 2014
Halorubrum rutilum Yin &al., 2016
Halorubrum saccharovorum (Tomlinson & Hochstein, 1977), McGenity & Grant, 1996
Halorubrum salinum Zhang & Cui, 2014
Halorubrum salsamenti Chen &al., 2019
Halorubrum sodomense (Oren, 1983) McGenity & Grant, 1996
Halorubrum tebenquichense Lizama &al., 2002
Halorubrum terrestre Ventosa &al., 2004
Halorubrum tibetense Fan &al., 2004
Halorubrum trapanicum (Petter, 1931) McGenity & Grant, 1996
Halorubrum trueperi Chen &al., 2017
††‡ Halorubrum vacuolatum (Mwatha & Grant, 1993) Kamekura &al., 1997; Oren, 2002
Halorubrum xinjiangense Fen &al., 2016
Halorubrum yunnanense Chen &al., 2015
Halorussus amylolyticus Yuan &al., 2015
Halorussus halophilus Ding &al., 2020
Halorussus litoreus Han &al., 2019
Halorussus rarus Cui &al., 2014
Halorussus ruber Xu &al., 2015
Halorussus salinus Xu &al., 2017
Halosimplex carlsbadense Vreeland &al., 2003
Halosimplex litoreum Yuan &al., 2015
Halosimplex pelagicum Han & Cui, 2014
Halosimplex rubrum Han & Cui, 2014
Halostagnicola alkaliphila Nagaoka &al., 2011
Halostagnicola bangensis Corral &al., 2015
Halostagnicola kamekurae Nagaoka &al., 2010
Halostagnicola larsenii Castillo &al., 2006
Haloterrigena daquingensis Wang &al., 2010
Haloterrigena hispanica Romano &al., 2007
Haloterrigena jeotgali Roh &al., 2009
Haloterrigena limicola Cui &al., 2006
Haloterrigena long Cui &al., 2006
Haloterrigena mahii Ding &al., 2017
Haloterrigena saccharevitans Xu &al., 2005
Haloterrigena salifodinae Chen &al., 2020
Haloterrigena salina Gutierrez &al., 2008
Haloterrigena thermotolerans Montalvo-Rodriguez &al., 2000
Haloterrigena turkmenica (Zvyagintseva & Tarasov, 1989) Ventosa &al., 1999
Halovivax asiaticus Castillo &al., 2006
Halovivax cerinus Amoozegar &al., 2015
Halovivax limisalsi Amoozegar &al., 2014
Halovivax ruber Castillo &al., 2007
Candidatus Iainarchaeum andersonii Rinke, &al., 2003; Baker &al., 2006
Candidatus Mancarchaeum acidiphilum Baker &al., 2006; Golyshina &al., 2017
Metallosphaera cuprina Zhang &al., 2015
Metallosphaera hakonensis Zhang &al., 2015
Metallosphaera prunae Johnson, 1998; Zhang &al., 2015
Metallosphaera sedula Johnson, 1998; Fashola &al., 2015; Zhang &al., 2015
Metallosphaera yellowstonensis Zhang &al., 2015
Methanocaldocococcus jannaschii
(Methanococcus jannaschii)
Brock &al., 1972
* Methanocaldocococcus thermolithotrophicus Brock &al., 1972
* Methanococcoides Morita & Moyer, 2001
* Methanogenium Morita & Moyer, 2001
* Methanosarcina Morita & Moyer, 2001
Methanopyrus kandleri Kurr &al., 1992
* Methanosarcina Morita & Moyer, 2001
Candidatus Micrarchaeum acidiphilum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Nanoarchaeum equitans Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Nanobsidianus stetteri
Nanopusillus acidilobi sp.
Candidatus Nanosalina sp. Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Nanosalinarum sp. Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Natrialba aegyptica Hezayen &al., 2001
Natrialba asiatica
Natrialba chahannaoensis Xu &al., 2001
Natrialba hulunbeirensis Xu &al., 2001
***††† Natrialba magadii (Tindall &al., 1984) Kamekura &al., 1997
Natrialba swarupiae Kajale &al., 2020
Natrialba taiwanensis/i> Hezayen &al., 2001; Oren, 2002
Natrinema altunense Xu &al., 2005
Natrinema ejinorense Castillo &al., 2006
Natrinema gari Tapingkae &al., 2008
Natrinema pallidum McGenity &al., 1998
Natrinema salaciae Albuquerque &al., 2012
Natrinema soli Rasooli &al., 2017
Natrinema versiforme Xin &al., 2000
Natrarchaeobaculum aegyptiacum (Zhao &al., 2018) Sorokin &al., 2020
Natronoarchaeum sulfireducens (Sorokin &al., 2019) Sorokin &al., 2020
Natronoarchaeum persicum Naghoni &al., 2017
Natronoarchaeum philippinense Shimane &al., 2013
Natronoarchaeum rubrumm Qiu &al., 2014
††‡ Natronobacterium gregoryi Tindall &al. 1984; Oren, 2002
Natronobacterium texcoconense Ruiz-Romero &al. 2013
Natronococcus amylolyticus Kanai &al. 1995
Natronococcus jeotgali Roh &al. 2007
††‡ Natronococcus occultus Tindall &al. 1984
Natronococcus roseus Corral &al. 2013
Natrolimnobius baerhuensis Itoh &al. 2005
Natrolimnohabitans innermongolicus Itoh &al. 2005
Natronomonas gomsonensis Kim &al. 2014
Natronomonas halophila Yin &al. 2020
Natronomonas moolapensis Burns &al. 2010
††‡ Natronomonas pharaonis (Soliman & Truper, 1983) Kamekura &al. 1997
Natronomonas salina Yin &al. 2020
Natronomonas salsuginis Duran-Viseras &al. 2020
Natronorubrum aibiense Cui &al. 2006
Natronorubrum bangense Xu &al. 1999
Natronorubrum halophilum Tao &al. 2020
Natronorubrum sediminis Gutierrez &al. 2010
Natronorubrum sulfidifaciens Cui &al. 2007
Natronorubrum texcoconense Ruiz-Romero &al. 2013
Natronorubrum tibetense Xu &al. 1999
Candidatus Nitrosoarchaeum koreensis Herbold &al., 2017
Candidatus Nitrosoarchaeum limnia Herbold &al., 2017
Candidatus Nitrosoarchaeum limnia Herbold &al., 2017
Candidatus Nitrosocosmicus oleophilus Herbold &al., 2017
Candidatus Nitrosopelagicus brevis Herbold &al., 2017
Candidatus Nitrosopumilus maritimus Lehtovirta-Morley &al., 2014; Herbold &al., 2017
Candidatus Nitrosopumilus viennensis Lehtovirta-Morley &al., 2014
Candidatus Nitrosopumilus sp. SJ Herbold &al., 2017
Candidatus Nitrosopumilus sp. NF5 Herbold &al., 2017
Candidatus Nitrosopumilus sp. AR1 Herbold &al., 2017
Candidatus Nitrosopumilus sp. AR2 Herbold &al., 2017
Candidatus Nitrosopumilus salaria Herbold &al., 2017
Candidatus Nitrosopumilus sp. D3C Herbold &al., 2017
Candidatus Nitrososphaera evergladensis Herbold &al., 2017
Candidatus Nitrososphaera viennensis Herbold &al., 2017
Candidatus Nitrososphaera gargensis Herbold &al., 2017
Candidatus Nitrosotalea okcheonensis Herbold &al., 2017
Candidatus Nitrosotalea sinensis Herbold &al., 2017
Candidatus Nitrosotalea devanaterra Lehtovirta-Morley &al., 2011 & 2014; Herbold &al., 2017
Candidatus Nitrosotalea bavarica Herbold &al., 2017
Candidatus Nitrosotenuis uzonensis Herbold &al., 2017
Candidatus Nitrosotenuis chungbukensis Herbold &al., 2017
Candidatus Nitrosotenuis cloacae Herbold &al., 2017
Candidatus Parvarchaeum acidiphilum Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Candidatus Parvarchaeum acidophilus Baker &al., 2006; Baker &al., 2010; Golyshina &al., 2017
Picrophilus oshimae Rampelotto, 2013, Johnson, 1998
Picrophilus torridus Rampelotto, 2013, Johnson, 1998; Zhang &al., 2015
Picrophilus sp. (Baker-Austin & Dopson, 2007)
Pyrococcus abyssi Erauso &al., 1993
Pyrococcus endeavori ?
Pyrococcus furiosus Erauso &al., 1993
Pyrococcus glycovorans ?
Pyrococcus horikoshii Erauso &al., 1993
Pyrococcus woesei ?
Pyrolobus fumarii Bloch &al., 1997
Salarchaeum japonicum Shimane &al. 2011
Stygiolobus azoricus Johnson, 1998; Zhang &al., 2015; Segerer &al., 1991
Sulfolobus acidocaldarius Johnson, 1998; Edwards &al., 2000; Zhang &al., 2015; Brock &al., 1972
Sulfolobus hakonensis Johnson, 1998
Sulfolobus islandicus Brock &al., 1972
Sulfolobus metallicus Johnson, 1998; Fashola &al., 2015; Zhang &al., 2015; Brock &al., 1972
Sulfolobus mirabilis Johnson, 1998
Sulfolobus neozealandicus Brock &al., 1972
Sulfolobus rivotinct Zhang &al., 2015
Sulfolobus shibatae Johnson, 1998; Zhang &al., 2015; Brock &al., 1972
Sulfolobus solfataricus Johnson, 1998, Dopson (2003); Zhang &al., 2015; Brock &al., 1972
Sulfolobus tengchongensis Zhang &al., 2015; Brock &al., 1972
Sulfolobus thuringiensis Brock &al., 1972
Sulfolobus tokodaii Dopson 2003; Zhang &al., 2015; Brock &al., 1972
Sulfolobus yangmingensis Zhang &al., 2015; Brock &al., 1972
Sulfolobus sp. AMP12/99 ?
Sulfolobus sp. CH7/99 ?
Sulfolobus sp. FF5/00 ?
Sulfolobus sp. G81 ?
Sulfolobus sp. H ?
Sulfolobus sp. JP2 ?
Sulfolobus sp. JP3 ?
Sulfolobus sp. K00 8-41 ?
Sulfolobus sp. L00 11 ?
Sulfolobus sp. L00 24 ?
Sulfolobus sp. M02 ?
Sulfolobus sp. MV2/99 ?
Sulfolobus sp. MVSoil3/SC2 ?
Sulfolobus sp. MVSoil6/SC1 ?
Sulfolobus sp. NGB6/00 ?
Sulfolobus sp. NGB23/00 ?
Sulfolobus sp. NGL8/00 ?
Sulfolobus sp. NO82 ?
Sulfolobus sp. NOB8H2 ?
Sulfolobus sp. RC3 ?
Sulfolobus sp. RC6/00 ?
Sulfolobus sp. RCSC1/01 ?
Sulfolobus sp. Sko-3 ?
Sulfolobus sp. ?
Sulfolobus sp. Ta ?
Sulfolobus sp. Tu A ?
Sulfolobus sp. Tu B-1 ?
Sulfolobus sp. Y00 58-82 ?
Sulfolobus sp. Y01 90#-18 ?
Sulfolobus sp. Y99 9-16 ?
Sulfolobus sp. Y99 9-23 ?
Sulfolobus sp. (uncultured) ?
Sulfurisphaera ohwakuensis Zhang &al., 2015
Sulfurococcus mirabilis Zhang &al., 2015
Sulfurococcus yellowstonii Johnson, 1998
Sulfurococcus yellowstonensis Zhang &al., 2015
* Thermococcus acidaminovorans ?
* Thermococcus aegaeus ?
* Thermococcus aggregans ?
*†† Thermococcus alcaliphilus Keller &al., 1997
* Thermococcus atlanticus ?
*** Thermococcus barophilus Marteinsson &al., 1999
* Thermococcus barossi ?
*† Thermococcus celer Zillig, 1983
*** Thermococcus chitonophagus
(one of only three archaeal chiton-philes)
Huber & Stetter, 1996
* Thermococcus coalescens ?
* Thermococcus fumicolans ?
* Thermococcus gammatolerans Jolivet, 2003
* Thermococcus gorgonarius ?
* Thermococcus guaymasensis ?
*** Thermococcus hydrothermalis Godfroy &al., 1997
* Thermococcus kodakarensis Atomi &al., 2004
***† Thermococcus litoralis Neuner &al., 2001
* Thermococcus marinus ?
* Thermococcus mexicalis ?
* Thermococcus pacificus ?
*** Thermococcus peptonophilus (more thermophilic at higher pressures and more barophilic at higher temps Canganella &al., 1996
* Thermococcus profundis Kobayashi & Horikoshi, 1995
* Thermococcus radiotolerans ?
* Thermococcus sibiricus ?
* Thermococcus siculi ?
* Thermococcus stetteri Miroshnichenko, 1990
* Thermococcus waimanguensis ?
* Thermococcus waiotapuensis ?
*††? Thermococcus zilligii Zillig, 1982
* Thermococcus sp. ?
* Thermococcus sp. 12-4 NCBI’s Taxonomy Browser
* Thermococcus sp. 13-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 13-3 NCBI’s Taxonomy Browser
* Thermococcus sp. 21-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 23-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 23-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 26-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 26-3 NCBI’s Taxonomy Browser
* Thermococcus sp. 28-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 29-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 300-Tc NCBI’s Taxonomy Browser
* Thermococcus sp. 30-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 31-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 31-3 NCBI’s Taxonomy Browser
* Thermococcus sp. 5-1 NCBI’s Taxonomy Browser
* Thermococcus sp. 70-4-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 83-5-2 NCBI’s Taxonomy Browser
* Thermococcus sp. 9N2 NCBI’s Taxonomy Browser
* Thermococcus sp. 9N3 NCBI’s Taxonomy Browser
* Thermococcus sp. 9oN-7 NCBI’s Taxonomy Browser
* Thermococcus sp. AEPII 1a NCBI’s Taxonomy Browser
* Thermococcus sp. AM4 NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete70-I78 NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete70-SCI NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete85-I78 NCBI’s Taxonomy Browser
* Thermococcus sp. Anhete 85-SCI NCBI’s Taxonomy Browser
* Thermococcus sp. AT1273 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-17 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-27 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-39 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax00-45 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-2 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-37 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-39 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-3 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-61 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-62 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax01-65 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax98-43 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax98-46 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax98-48 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax99-47 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax99-57 NCBI’s Taxonomy Browser
* Thermococcus sp. Ax99-67 NCBI’s Taxonomy Browser
* Thermococcus sp. B1001 NCBI’s Taxonomy Browser
* Thermococcus sp. BHI60a21 NCBI’s Taxonomy Browser
* Thermococcus sp. BHI80a28 NCBI’s Taxonomy Browser
* Thermococcus sp. BHI80a40 NCBI’s Taxonomy Browser
* Thermococcus sp. CAR-80 NCBI’s Taxonomy Browser
* Thermococcus sp. CKU-199 NCBI’s Taxonomy Browser
* Thermococcus sp. CL1 NCBI’s Taxonomy Browser
* Thermococcus sp. CL2 NCBI’s Taxonomy Browser
* Thermococcus sp. CMI NCBI’s Taxonomy Browser
* Thermococcus sp. CNR-5 NCBI’s Taxonomy Browser
* Thermococcus sp. CX1 NCBI’s Taxonomy Browser
* Thermococcus sp. CX2 NCBI’s Taxonomy Browser
* Thermococcus sp. CX3 NCBI’s Taxonomy Browser
* Thermococcus sp. CX4 NCBI’s Taxonomy Browser
* Thermococcus sp. CYA NCBI’s Taxonomy Browser
* Thermococcus sp. Dex80a71 NCBI’s Taxonomy Browser
* Thermococcus sp. Dex80a75 NCBI’s Taxonomy Browser
* Thermococcus sp. enrichment clone SA3 NCBI’s Taxonomy Browser
* Thermococcus sp. GE8 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda2 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda3 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda4 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda5 NCBI’s Taxonomy Browser
* Thermococcus sp. Gorda6 NCBI’s Taxonomy Browser
* Thermococcus sp. GT NCBI’s Taxonomy Browser
* Thermococcus sp. GU5L5 NCBI’s Taxonomy Browser
* Thermococcus sp. JDF-3 NCBI’s Taxonomy Browser
* Thermococcus sp. KI NCBI’s Taxonomy Browser
* Thermococcus sp. KS-1 NCBI’s Taxonomy Browser
* Thermococcus sp. KS-8 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1031 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1049 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1083 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1092 NCBI’s Taxonomy Browser
* Thermococcus sp. MV1099 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ10 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ11 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ12 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ13 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ1 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ2 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ3 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ5 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ6 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ8 NCBI’s Taxonomy Browser
* Thermococcus sp. MZ9 NCBI’s Taxonomy Browser
* Thermococcus sp. NA1 NCBI’s Taxonomy Browser
* Thermococcus sp. NS85-T NCBI’s Taxonomy Browser
* Thermococcus sp. OGL-20P NCBI’s Taxonomy Browser
* Thermococcus sp. P6 NCBI’s Taxonomy Browser
* Thermococcus sp. Pd70 NCBI’s Taxonomy Browser
* Thermococcus sp. Pd85 NCBI’s Taxonomy Browser
* Thermococcus sp. Rt3 NCBI’s Taxonomy Browser
* Thermococcus sp. SB611 NCBI’s Taxonomy Browser
* Thermococcus sp. SN531 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-1-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-1-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-1-95 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-2-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-2-95 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-365-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-365-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-365-95 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-4-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-4-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-CRC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-CRC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-MC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-SC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc70-SC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-0 age SC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-CRC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-CRC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-MC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-MC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-SC-ISCS NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-SC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc85-SC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-CRC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-CRC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-MC-I NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-MC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc95-SC-S NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-I-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-I-85 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-S-70 NCBI’s Taxonomy Browser
* Thermococcus sp. Tc-S-85 NCBI’s Taxonomy Browser
* Thermococcus sp. TK1 NCBI’s Taxonomy Browser
* Thermococcus sp. TM1 NCBI’s Taxonomy Browser
* Thermococcus sp. TS2 NCBI’s Taxonomy Browser
* Thermococcus sp. TS3 NCBI’s Taxonomy Browser
* Thermococcus sp. vp197 NCBI’s Taxonomy Browser
* Thermococcus sp. (uncultured) ?
Thermogymnomonas acidocola Zhang &al., 2015
Thermoplasma acidophilum Johnson, 1998; Zhang &al., 2015
Thermoplasma volcanium Johnson, 1998; Zhang &al., 2015



Full Citations

Schuerger & Nicholson, 2016.
http://doi.org/10.1089/ast.2016.1587

Nicholson &al., 2013.
https://doi.org/10.1073/pnas.1209793110

Golyshina, O.V.; Yakimov, M.M.; Lunsdorf, H.; Ferrer, M; Nimtz, M.; Timmis, K.N.; Wray, V.; Tindall, B.J.; Golyshin, P.N.; (2009); Acidiplasma aeolicum gen. nov., sp. nov., a euryarchaeon of the family Ferroplasmaceae isolated from a hydrothermal pool, and transfer of Ferroplasma cupricumulans to Acidiplasma cupricumulans comb. nov



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